{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,9]],"date-time":"2026-10-09T18:56:47Z","timestamp":1791572207414,"version":"4.3.4"},"reference-count":81,"publisher":"MDPI AG","issue":"8","license":[{"start":{"date-parts":[[2024,8,6]],"date-time":"2024-08-06T00:00:00Z","timestamp":1722902400000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"name":"\u201cRegional Excellence Initiative\u201d Program for 2024\u20132027","award":["RID\/SP\/0045\/2024\/01"],"award-info":[{"award-number":["RID\/SP\/0045\/2024\/01"]}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Pathogens"],"abstract":"<jats:p>Rickettsiae are an interesting group of bacteria comprising a large number of obligate intracellular species. The circulation of these bacteria in the environment depends on the presence of vectors (blood-sucking invertebrates) and their hosts. On the basis of phylogenetic analyses in 2022, a division into five groups of Rickettsia has been proposed: I belli group, II canadensis group, III typhus group, and IV and V spotted group fever (respectively II, phylogenetically older, and I). The genus Rickettsia includes species that are both pathogenic and nonpathogenic to humans and domestic and wild animals. Some Rickettsia species are invertebrate symbionts. Currently, rickettsiae, which are transmitted mainly by ticks, are spreading worldwide. This has been promoted by climate change, environmental changes caused by humans, and the synanthropisation of plants and animals. Therefore, it is extremely important to monitor the natural and urban environments. The study of potential vectors and reservoirs of bacteria in the genus Rickettsia should be a permanent part of the analysis of the modern human environment.<\/jats:p>","DOI":"10.3390\/pathogens13080661","type":"journal-article","created":{"date-parts":[[2024,8,6]],"date-time":"2024-08-06T15:24:16Z","timestamp":1722957856000},"page":"661","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":33,"title":["Rickettsia Species: Genetic Variability, Vectors, and Rickettsiosis\u2014A Review"],"prefix":"10.3390","volume":"13","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-1453-3006","authenticated-orcid":false,"given":"Anna","family":"Rymaszewska","sequence":"first","affiliation":[{"name":"Department of Genetics and Genomics, Institute of Biology, University of Szczecin, ul. Felczaka 3C, 71-412 Szczecin, Poland"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-4386-7766","authenticated-orcid":false,"given":"Mariusz","family":"Piotrowski","sequence":"additional","affiliation":[{"name":"Department of Genetics and Genomics, Institute of Biology, University of Szczecin, ul. Felczaka 3C, 71-412 Szczecin, Poland"},{"name":"BIOSPACE Foundation, ul. Karpia 31, 61-619 Pozna\u0144, Poland"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2024,8,6]]},"reference":[{"key":"ref_1","doi-asserted-by":"crossref","first-page":"126","DOI":"10.1016\/j.ijid.2014.05.025","article-title":"Syndromic classification of rickettsioses: An approach for clinical practice","volume":"28","author":"Hidalgo","year":"2014","journal-title":"Int. J. Infect. Dis."},{"key":"ref_2","doi-asserted-by":"crossref","first-page":"2630","DOI":"10.1038\/s41467-022-30385-6","article-title":"Genomic diversity across the Rickettsia and \u2018Candidatus Megaira\u2019 genera and proposal of genus status for the Torix group","volume":"13","author":"Davison","year":"2022","journal-title":"Nat. Commun."},{"key":"ref_3","doi-asserted-by":"crossref","unstructured":"Chaorattanakawee, S., Korkusol, A., Tippayachai, B., Promsathaporn, S., Poole-Smith, B.K., and Takhampunya, R. (2021). Amplicon-Based Next Generation Sequencing for rapid identification of Rickettsia and ectoparasite species from entomological surveillance in Thailand. Pathogens, 10.","DOI":"10.3390\/pathogens10020215"},{"key":"ref_4","doi-asserted-by":"crossref","unstructured":"Stewart, A.G., and Stewart, A.G.A. (2021). An update on the laboratory diagnosis of Rickettsia spp. infection. Pathogens, 10.","DOI":"10.3390\/pathogens10101319"},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"379","DOI":"10.1111\/j.1469-185X.2010.00151.x","article-title":"Rickettsial evolution in the light of comparative genomics","volume":"86","author":"Merhej","year":"2011","journal-title":"Biol. Rev."},{"key":"ref_6","doi-asserted-by":"crossref","unstructured":"Piotrowski, M., and Rymaszewska, A. (2020). Expansion of Tick-Borne Rickettsioses in the World. Microorganisms, 8.","DOI":"10.3390\/microorganisms8121906"},{"key":"ref_7","first-page":"1043","article-title":"Report of the ad hoc committee for the re-evaluation of the species definition in bacteriology","volume":"52","author":"Stackebrandt","year":"2002","journal-title":"Int. J. Syst. Evol. Microbiol."},{"key":"ref_8","doi-asserted-by":"crossref","first-page":"5456","DOI":"10.1128\/JCM.41.12.5456-5465.2003","article-title":"Gene sequence-based criteria for identification of new rickettsia isolates and description of Rickettsia heilongjiangensis sp. nov","volume":"41","author":"Fournier","year":"2003","journal-title":"J. Clin. Microbiol."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"1738","DOI":"10.1099\/ijsem.0.003963","article-title":"Genome sequence-based criteria for demarcation and definition of species in the genus Rickettsia","volume":"70","author":"Diop","year":"2020","journal-title":"Int. J. Syst. Evol. Microbiol."},{"key":"ref_10","doi-asserted-by":"crossref","unstructured":"Gillespie, J.J., Williams, K., Shukla, M., Snyder, E.E., Nordberg, E.K., Ceraul, S.M., Dharmanolla, C., Rainey, D., Soneja, J., and Shallom, J.M. (2008). Rickettsia phylogenomics: Unwinding the intricacies of obligate intracellular life. PLoS ONE, 3.","DOI":"10.1371\/journal.pone.0002018"},{"key":"ref_11","doi-asserted-by":"crossref","unstructured":"Gillespie, J.J., Beier, M.S., Rahman, M.S., Ammerman, N.C., Shallom, J.M., Purkayastha, A., Sobral, B.S., and Azad, A.F. (2007). Plasmids and rickettsial evolution: Insight from Rickettsia felis. PLoS ONE, 2.","DOI":"10.1371\/journal.pone.0000266"},{"key":"ref_12","doi-asserted-by":"crossref","first-page":"657","DOI":"10.1128\/CMR.00032-13","article-title":"Update on tick-borne rickettsioses around the world: A geographic approach","volume":"26","author":"Parola","year":"2013","journal-title":"Clin. Microbiol. Rev."},{"key":"ref_13","doi-asserted-by":"crossref","unstructured":"El Karkouri, K., Ghigo, E., Raoult, D., and Fournier, P.E. (2022). Genomic evolution and adaptation of arthropod-associated Rickettsia. Sci. Rep., 12.","DOI":"10.1038\/s41598-022-07725-z"},{"key":"ref_14","doi-asserted-by":"crossref","unstructured":"Shi, M., Qin, T., Liu, Z., Feng, H., Sun, Y., and Zhang, M. (2022). Meta-transcriptional detection of Rickettsia canadensis from Ixodes persulcatus in China. Res. Sq.","DOI":"10.21203\/rs.3.rs-2239865\/v1"},{"key":"ref_15","doi-asserted-by":"crossref","first-page":"94","DOI":"10.1099\/00207713-33-1-94","article-title":"Rickettsia bellii sp. nov.: A tick-borne Rickettsia, widely distributed in the United States, that is distinct from the spotted fever and typhus biogroups","volume":"33","author":"Philip","year":"1983","journal-title":"Int. J. Syst. Bacteriol."},{"key":"ref_16","doi-asserted-by":"crossref","first-page":"798","DOI":"10.1099\/00207713-44-4-798","article-title":"Ancestral divergence of Rickettsia bellii from the spotted fever and typhus groups of Rickettsia and antiquity of the genus Rickettsia","volume":"44","author":"Stothard","year":"1994","journal-title":"Int. J. Syst. Bacteriol."},{"key":"ref_17","doi-asserted-by":"crossref","unstructured":"Krawczak, F.S., Labruna, M.B., Hecht, J.A., Paddock, C.D., and Karpathy, S.E. (2018). Genotypic characterization of Rickettsia bellii reveals distinct lineages in the United States and South America. BioMed Res. Int., 2018.","DOI":"10.1155\/2018\/8505483"},{"key":"ref_18","doi-asserted-by":"crossref","first-page":"482","DOI":"10.1016\/j.ttbdis.2016.02.004","article-title":"\u2018Candidatus Rickettsia mendelii\u2019, a novel basal group rickettsia detected in Ixodes ricinus ticks in the Czech Republic","volume":"7","author":"Hajduskova","year":"2016","journal-title":"Ticks Tick-Borne Dis."},{"key":"ref_19","doi-asserted-by":"crossref","first-page":"102104","DOI":"10.1016\/j.ttbdis.2022.102104","article-title":"Multilocus genetic analysis indicates taxonomic status of \u201cCandidatus Rickettsia mendelii\u201d as a separate basal group","volume":"14","author":"Igolkina","year":"2023","journal-title":"Ticks Tick-Borne Dis."},{"key":"ref_20","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1023\/A:1025381712339","article-title":"The Argasidae, Ixodidae and Nuttalliellidae (Acari: Ixodida): A world list of valid tick names","volume":"28","author":"Horak","year":"2002","journal-title":"Exp. Appl. Acarol."},{"key":"ref_21","doi-asserted-by":"crossref","first-page":"1","DOI":"10.11646\/zootaxa.3767.1.1","article-title":"Names for Ixodidae (Acari: Ixodoidea): Valid, synonyms, incertae sedis, nomina dubia, nomina nuda, lapsus, incorrect and suppressed names\u2014With notes on confusions and misidentifications","volume":"3767","author":"Guglielmone","year":"2014","journal-title":"Zootaxa"},{"key":"ref_22","doi-asserted-by":"crossref","first-page":"1","DOI":"10.11646\/zootaxa.2528.1.1","article-title":"The Agrasidae, Ixodidae and Nuttalliedae (Acari: Ixodida) of the world: A list of valid species names","volume":"2528","author":"Guglielmone","year":"2010","journal-title":"Zootaxa"},{"key":"ref_23","doi-asserted-by":"crossref","first-page":"105527","DOI":"10.1016\/j.actatropica.2020.105527","article-title":"Ticks and tick-borne pathogens in animals and humans in the island nations of Southeast Asia: A review","volume":"209","author":"Sharifah","year":"2020","journal-title":"Acta Trop."},{"key":"ref_24","doi-asserted-by":"crossref","first-page":"104","DOI":"10.1016\/j.antiviral.2014.05.016","article-title":"The ecology of ticks and epidemiology of tick-borne viral diseases","volume":"108","year":"2014","journal-title":"Antiviral. Res."},{"key":"ref_25","doi-asserted-by":"crossref","unstructured":"Gray, J.S., Kahl, O., Lane, R.S., and Stanek, G. (2002). Ecological research on Borrelia burgdorferi sensu lato: Terminology and some methodological pitfalls. Lyme Borreliosis Biology Epidemiology and Control (2002) CABI Books, CABI International.","DOI":"10.1079\/9780851996325.0000"},{"key":"ref_26","doi-asserted-by":"crossref","first-page":"548","DOI":"10.1099\/00207713-47-2-548","article-title":"Rickettsia aeschlimannii sp. nov., a new spotted fever group rickettsia associated with Hyalomma marginatum ticks","volume":"47","author":"Beati","year":"1997","journal-title":"Int. J. Syst. Bacteriol."},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"5569","DOI":"10.1128\/AEM.00122-06","article-title":"Isolation and identification of Rickettsia massiliae from Rhipicephalus sanguineus ticks collected in Arizona","volume":"72","author":"Eremeeva","year":"2006","journal-title":"Appl. Environ. Microbiol."},{"key":"ref_28","doi-asserted-by":"crossref","first-page":"122","DOI":"10.1016\/j.meegid.2014.03.014","article-title":"Genotyping, evolution and epidemiological findings of Rickettsia species","volume":"25","author":"Merhej","year":"2014","journal-title":"Infect. Genet. Evol."},{"key":"ref_29","doi-asserted-by":"crossref","first-page":"809","DOI":"10.1603\/0022-2585-39.6.809","article-title":"Rickettsial infection in Dermacentor variabilis (Acari: Ixodidae) inhibits transovarial transmission of a second Rickettsia","volume":"39","author":"Macaluso","year":"2002","journal-title":"J. Med. Entomol."},{"key":"ref_30","doi-asserted-by":"crossref","first-page":"289","DOI":"10.1093\/jmedent\/45.2.289","article-title":"Infection and co-infection rates of Anaplasma phagocytophilum variants, Babesia spp., Borrelia burgdorferi, and the rickettsial endosymbiont in Ixodes scapularis (Acari: Ixodidae) from sites in Indiana, Maine, Pennsylvania, and Wisconsin","volume":"45","author":"Steiner","year":"2008","journal-title":"J. Med. Entomol."},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"999","DOI":"10.1128\/AEM.68.2.999-1004.2002","article-title":"Novel clade of Rickettsia spp. from leeches","volume":"68","author":"Kikuchi","year":"2002","journal-title":"Appl. Environ. Microbiol."},{"key":"ref_32","first-page":"2097","article-title":"The emerging diversity of Rickettsia","volume":"273","author":"Perlman","year":"2006","journal-title":"Proc. Biol. Sci."},{"key":"ref_33","doi-asserted-by":"crossref","first-page":"388","DOI":"10.1128\/jb.176.2.388-394.1994","article-title":"Rickettsial relative associated with male killing in the ladybird beetle (Adalia bipunctata)","volume":"176","author":"Werren","year":"1994","journal-title":"J. Bacteriol."},{"key":"ref_34","first-page":"1791","article-title":"Horizontal transmission of the insect symbiont Rickettsia is plant-mediated","volume":"279","author":"Inbar","year":"2012","journal-title":"Proc. Biol. Sci."},{"key":"ref_35","doi-asserted-by":"crossref","first-page":"155","DOI":"10.1603\/ME12086","article-title":"Effects of infection by Arsenophonus and Rickettsia bacteria on the locomotive ability of the ticks Amblyomma americanum, Dermacentor variabilis, and Ixodes scapularis","volume":"50","author":"Kagemann","year":"2013","journal-title":"J. Med. Entomol."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"638","DOI":"10.1128\/iai.27.2.638-642.1980","article-title":"Sexual transmission of spotted fever group rickettsiae by infected male ticks: Detection of rickettsiae in immature spermatozoa of Ixodes ricinus","volume":"27","author":"Hayes","year":"1980","journal-title":"Infect. Immun."},{"key":"ref_37","doi-asserted-by":"crossref","first-page":"897","DOI":"10.1086\/319347","article-title":"Ticks and Tickborne Bacterial Diseases in Humans: An Emerging Infectious Threat","volume":"32","author":"Parola","year":"2001","journal-title":"Clin. Infect. Dis."},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"176","DOI":"10.1186\/s13071-020-04049-7","article-title":"Transovarial transmission of Borrelia spp., Rickettsia spp. and Anaplasma phagocytophilum in Ixodes ricinus under field conditions extrapolated from DNA detection in questing larvae","volume":"13","author":"Hauck","year":"2020","journal-title":"Parasites Vectors"},{"key":"ref_39","doi-asserted-by":"crossref","unstructured":"Wechtaisong, W., Bonnet, S.I., Chomel, B.B., Lien, Y.Y., Chuang, S.T., and Tsai, Y.L. (2021). Investigation of transovarial transmission of Bartonella henselae in Rhipicephalus sanguineus sensu lato ticks using artificial feeding. Microorganisms, 9.","DOI":"10.3390\/microorganisms9122501"},{"key":"ref_40","doi-asserted-by":"crossref","first-page":"259","DOI":"10.26444\/aaem\/167427","article-title":"Prevalence of pathogens in sympatric Ixodes ricinus and Dermacentor reticulatus ticks in Eastern Poland and their potential impact on oral-anal contacts between ticks","volume":"30","author":"Buczek","year":"2023","journal-title":"Ann. Agric. Environ. Med."},{"key":"ref_41","doi-asserted-by":"crossref","first-page":"686","DOI":"10.26444\/aaem\/99054","article-title":"A unique phenomenon of oral-anal contact between ticks observed in two tick species Ixodes ricinus and Dermacentor reticulatus","volume":"25","author":"Buczek","year":"2018","journal-title":"Ann. Agric. Environ. Med."},{"key":"ref_42","doi-asserted-by":"crossref","first-page":"1752","DOI":"10.1080\/21505594.2022.2132047","article-title":"Pathogenicity and virulence of Rickettsia","volume":"13","author":"Helminiak","year":"2022","journal-title":"Virulence"},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"755","DOI":"10.1023\/A:1018473122070","article-title":"Infestation of mammals by Ixodes ricinus ticks (Acari: Ixodidae) in south-central Sweden","volume":"21","author":"Talleklint","year":"1997","journal-title":"Exp. Appl. Acarol."},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"75","DOI":"10.1186\/s13071-022-05641-9","article-title":"Worldwide host associations of the tick genus Ixodes suggest relationships based on environmental sharing rather than on co-phylogenetic events","volume":"16","author":"Guglielmone","year":"2023","journal-title":"Parasit. Vec."},{"key":"ref_45","doi-asserted-by":"crossref","first-page":"179","DOI":"10.3201\/eid0402.980205","article-title":"Rickettsial Pathogens and Their Arthropod Vectors","volume":"4","author":"Azad","year":"1998","journal-title":"Emerg. Infect. Dis."},{"key":"ref_46","doi-asserted-by":"crossref","first-page":"149","DOI":"10.1186\/1756-3305-6-149","article-title":"Dissemination of bloodmeal acquired Rickettsia felis in cat fleas, Ctenocephalides felis","volume":"6","author":"Thepparit","year":"2013","journal-title":"Parasites Vectors"},{"key":"ref_47","doi-asserted-by":"crossref","first-page":"90","DOI":"10.1016\/j.ijpddr.2017.01.004","article-title":"Blocking transmission of vector-borne diseases","volume":"7","author":"Noack","year":"2017","journal-title":"Int. J. Parasitol. Drugs Drug Resist."},{"key":"ref_48","doi-asserted-by":"crossref","first-page":"452","DOI":"10.1186\/1756-3305-7-452","article-title":"Immediate efficacy and persistent speed of kill of a novel oral formulation of afoxolaner (NexGardTM) against induced infestations with Ixodes ricinus ticks","volume":"7","author":"Halos","year":"2014","journal-title":"Parasites Vectors"},{"key":"ref_49","doi-asserted-by":"crossref","first-page":"2","DOI":"10.1051\/parasite\/2016002","article-title":"Assessment of the prophylactic speed of kill of Frontline Tri-Act(\u00ae) against ticks (Ixodes ricinus and Rhipicephalus sanguineus) on dogs","volume":"23","author":"Beugnet","year":"2016","journal-title":"Parasite"},{"key":"ref_50","doi-asserted-by":"crossref","first-page":"37","DOI":"10.1016\/j.vetpar.2016.02.014","article-title":"Evaluation of the speed of kill of sarolaner (Simparica\u2122) against induced infestations of three species of ticks (Amblyomma maculatum, Ixodes scapularis, Ixodes ricinus) on dogs","volume":"30","author":"Six","year":"2016","journal-title":"Vet. Parasitol."},{"key":"ref_51","doi-asserted-by":"crossref","first-page":"781","DOI":"10.1099\/jmm.0.001206","article-title":"Emerging tick-borne pathogens of public health importance: A mini-review","volume":"69","author":"Rochlin","year":"2020","journal-title":"J. Med. Microbiol."},{"key":"ref_52","doi-asserted-by":"crossref","first-page":"719","DOI":"10.1128\/CMR.18.4.719-756.2005","article-title":"Tick-borne rickettsioses around the world: Emerging diseases challenging old concepts","volume":"18","author":"Parola","year":"2005","journal-title":"Clin. Microbiol. Rev."},{"key":"ref_53","unstructured":"(2024, April 22). Volunteer for Clinical Studies, Available online: https:\/\/www.niaid.nih.gov\/."},{"key":"ref_54","doi-asserted-by":"crossref","first-page":"180","DOI":"10.1016\/j.jpeds.2006.11.023","article-title":"Clinical and laboratory features, hospital course, and outcome of Rocky Mountain spotted fever in children","volume":"150","author":"Buckingham","year":"2007","journal-title":"J. Pediatr."},{"key":"ref_55","doi-asserted-by":"crossref","first-page":"793","DOI":"10.1590\/S0074-02762007000700003","article-title":"Rickettsia infection in five areas of the state of S\u00e3o Paulo, Brazil","volume":"102","author":"Horta","year":"2007","journal-title":"Mem\u00f3rias Inst. Oswaldo Cruz"},{"key":"ref_56","unstructured":"(2024, February 09). Brazil Has 49 Cases of Spotted Fever in 2023, Six Deaths. Available online: https:\/\/agenciabrasil.ebc.com.br\/en\/saude\/noticia\/2023-06\/brazil-has-49-cases-spotted-fever-2023-six-deaths."},{"key":"ref_57","doi-asserted-by":"crossref","first-page":"815","DOI":"10.3201\/eid2604.191251","article-title":"Needlestick-Associated Rocky Mountain Spotted Fever, Brazil","volume":"26","author":"Adelino","year":"2020","journal-title":"Emerg. Infect. Dis."},{"key":"ref_58","doi-asserted-by":"crossref","first-page":"112","DOI":"10.1016\/S0140-6736(05)61814-4","article-title":"A new tick-transmitted disease due to Rickettsia slovaca","volume":"12","author":"Raoult","year":"1997","journal-title":"Lancet"},{"key":"ref_59","doi-asserted-by":"crossref","first-page":"327","DOI":"10.1111\/j.1198-743X.2004.00782.x","article-title":"Dermacentor-borne necrosis erythema and lymphadenopathy: Clinical and epidemiological features of a new tick-borne disease","volume":"10","author":"Oteo","year":"2004","journal-title":"Clin. Microbiol. Infect."},{"key":"ref_60","doi-asserted-by":"crossref","first-page":"521","DOI":"10.3201\/eid1203.050911","article-title":"Rickettsia slovaca infection, France","volume":"12","author":"Gouriet","year":"2006","journal-title":"Emerg. Infect. Dis."},{"key":"ref_61","doi-asserted-by":"crossref","first-page":"549","DOI":"10.1086\/650172","article-title":"Scalp eschar and neck lymphadenopathy caused by Bartonella henselae after tick bite","volume":"15","author":"Angelakis","year":"2010","journal-title":"Clin. Infect. Dis."},{"key":"ref_62","unstructured":"Rickettsial Diseases (2024, February 09). CDC Yellow Book 2024. Travel-Associated Infections & Diseases, Available online: https:\/\/wwwnc.cdc.gov\/travel\/yellowbook\/2024\/infections-diseases\/rickettsial-diseases."},{"key":"ref_63","doi-asserted-by":"crossref","first-page":"e00852","DOI":"10.1016\/j.idcr.2020.e00852","article-title":"Rickettsia typhi central nervous system infection","volume":"29","author":"Talhelm","year":"2020","journal-title":"IDCases"},{"key":"ref_64","unstructured":"(2024, February 09). Rickettsiosis Subcommittee Report to the Tick-Borne Disease Working Group, Available online: https:\/\/www.hhs.gov\/ash\/advisory-committees\/tickbornedisease\/reports\/rickettsiosis-subcomm-2020\/index.html."},{"key":"ref_65","doi-asserted-by":"crossref","first-page":"101855","DOI":"10.1016\/j.ttbdis.2021.101855","article-title":"Rickettsiosis subcommittee report to the tick-borne disease working group","volume":"13","author":"Walker","year":"2022","journal-title":"Ticks Tick-Borne Dis."},{"key":"ref_66","doi-asserted-by":"crossref","first-page":"587131","DOI":"10.1155\/2015\/587131","article-title":"Human Coinfection with Borrelia burgdorferi and Babesia microti in the United States","volume":"2015","author":"Knapp","year":"2015","journal-title":"J. Parasitol. Res."},{"key":"ref_67","doi-asserted-by":"crossref","first-page":"1301","DOI":"10.3934\/mbe.2017067","article-title":"Modeling co-infection of Ixodes tick-borne pathogens","volume":"14","author":"Lou","year":"2017","journal-title":"Math. Biosci. Eng."},{"key":"ref_68","doi-asserted-by":"crossref","first-page":"265","DOI":"10.1089\/153036602321653851","article-title":"Epidemiology and impact of coinfections acquired from Ixodes ticks","volume":"2","author":"Belongia","year":"2002","journal-title":"Vector Borne Zoonotic Dis."},{"key":"ref_69","doi-asserted-by":"crossref","first-page":"601","DOI":"10.1016\/j.ijpara.2006.01.014","article-title":"Anaplasma phagocytophilum-infected neutrophils enhance transmigration of Borrelia burgdorferi across the human blood brain barrier in vitro","volume":"36","author":"Nyarko","year":"2006","journal-title":"Int. J. Parasitol."},{"key":"ref_70","doi-asserted-by":"crossref","unstructured":"Boyer, P.H., Lenormand, C., Jaulhac, B., and Talagrand-Reboul, E. (2022). Human co-infections between Borrelia burgdorferi s.l. and other Ixodes-borne microorganisms: A systematic review. Pathogens, 11.","DOI":"10.3390\/pathogens11030282"},{"key":"ref_71","doi-asserted-by":"crossref","first-page":"347","DOI":"10.1186\/1756-3305-6-347","article-title":"Co-infection of Borrelia burgdorferi sensu lato and Rickettsia species in ticks and in an erythema migrans patient","volume":"6","author":"Sprong","year":"2013","journal-title":"Parasites Vectors"},{"key":"ref_72","doi-asserted-by":"crossref","first-page":"26","DOI":"10.5604\/12321966.1233893","article-title":"Prevalence of infections and co-infections with 6 pathogens in Dermacentor reticulatus ticks collected in eastern Poland","volume":"24","author":"Sawczyn","year":"2017","journal-title":"Ann. Agric. Environ. Med."},{"key":"ref_73","doi-asserted-by":"crossref","first-page":"18","DOI":"10.1016\/j.ttbdis.2017.10.014","article-title":"Association of Borrelia and Rickettsia spp. and bacterial loads in Ixodes ricinus ticks","volume":"9","author":"Raulf","year":"2018","journal-title":"Ticks Tick-Borne Dis."},{"key":"ref_74","doi-asserted-by":"crossref","unstructured":"Moutailler, S., Valiente Moro, C., Vaumourin, E., Michelet, L., Tran, F.H., Devillers, E., Cosson, J.F., Gasqui, P., Van, V.T., and Mavingui, P. (2016). Co-infection of ticks: The rule rather than the exception. PLoS Negl. Trop. Dis., 10.","DOI":"10.1371\/journal.pntd.0004539"},{"key":"ref_75","doi-asserted-by":"crossref","unstructured":"Rocha, S.C., Vel\u00e1squez, C.V., Aquib, A., Al-Nazal, A., and Parveen, N. (2022). Transmission cycle of tick-borne infections and co-infections, animal models and diseases. Pathogens, 11.","DOI":"10.3390\/pathogens11111309"},{"key":"ref_76","doi-asserted-by":"crossref","first-page":"161","DOI":"10.1016\/j.ttbdis.2013.10.002","article-title":"Rickettsial infection in Ixodes ricinus ticks in urban and natural habitats of Slovakia","volume":"5","year":"2014","journal-title":"Ticks Tick-Borne Dis."},{"key":"ref_77","doi-asserted-by":"crossref","first-page":"573","DOI":"10.1186\/s13071-017-2391-2","article-title":"Ticks and the city\u2014Are there any differences between city parks and natural forests in terms of tick abundance and prevalence of spirochaetes?","volume":"10","author":"Kowalec","year":"2017","journal-title":"Parasite Vectors"},{"key":"ref_78","doi-asserted-by":"crossref","first-page":"941","DOI":"10.1016\/j.scitotenv.2019.03.235","article-title":"Ticks and tick-borne diseases in the city: Role of landscape connectivity and green space characteristics in a metropolitan area","volume":"670","author":"Heylen","year":"2019","journal-title":"Sci. Total Environ."},{"key":"ref_79","doi-asserted-by":"crossref","first-page":"591","DOI":"10.1111\/1758-2229.13075","article-title":"New problems of environmental ecology: Ticks and tick-borne pathogens in city parks of Ukraine","volume":"14","author":"Kolomiiets","year":"2022","journal-title":"Environ. Microbiol. Rep."},{"key":"ref_80","doi-asserted-by":"crossref","first-page":"102305","DOI":"10.1016\/j.ttbdis.2023.102305","article-title":"Nanopore-based metagenomics reveal a new Rickettsia in Europe","volume":"15","author":"Ergunay","year":"2024","journal-title":"Ticks Tick-Borne Dis."},{"key":"ref_81","doi-asserted-by":"crossref","unstructured":"Kipp, E.J., Lindsey, L.L., Khoo, B., Faulk, C., Oliver, J.D., and Larsen, P.A. (2023). Metagenomic surveillance for bacterial tick-borne pathogens using nanopore adaptive sampling. Sci. Rep., 13.","DOI":"10.1038\/s41598-023-37134-9"}],"container-title":["Pathogens"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.mdpi.com\/2076-0817\/13\/8\/661\/pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,10,10]],"date-time":"2025-10-10T15:31:04Z","timestamp":1760110264000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.mdpi.com\/2076-0817\/13\/8\/661"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2024,8,6]]},"references-count":81,"journal-issue":{"issue":"8","published-online":{"date-parts":[[2024,8]]}},"alternative-id":["pathogens13080661"],"URL":"https:\/\/doi.org\/10.3390\/pathogens13080661","relation":{},"ISSN":["2076-0817"],"issn-type":[{"value":"2076-0817","type":"electronic"}],"subject":[],"published":{"date-parts":[[2024,8,6]]}}}