{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,14]],"date-time":"2026-08-14T04:30:38Z","timestamp":1786681838145,"version":"build-2736575974"},"reference-count":52,"publisher":"MDPI AG","issue":"16","license":[{"start":{"date-parts":[[2026,8,12]],"date-time":"2026-08-12T00:00:00Z","timestamp":1786492800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100018542","name":"Natural Science Foundation of Sichuan Province","doi-asserted-by":"publisher","award":["2024NSFSC1329"],"award-info":[{"award-number":["2024NSFSC1329"]}],"id":[{"id":"10.13039\/501100018542","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100012453","name":"China Agricultural Research System","doi-asserted-by":"crossref","award":["CARS-10"],"award-info":[{"award-number":["CARS-10"]}],"id":[{"id":"10.13039\/501100012453","id-type":"DOI","asserted-by":"crossref"}]},{"award":["CARS-10"],"award-info":[{"award-number":["CARS-10"]}],"id":[{"id":"https:\/\/ror.org\/03p5eky76","id-type":"ROR","asserted-by":"publisher"}]},{"name":"Sichuan Academy of Agricultural Sciences","award":["1+9KJGG002"],"award-info":[{"award-number":["1+9KJGG002"]}]},{"award":["1+9KJGG002"],"award-info":[{"award-number":["1+9KJGG002"]}],"id":[{"id":"https:\/\/ror.org\/05f0php28","id-type":"ROR","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Plants"],"abstract":"<jats:p>Potassium (K+) is an essential mineral element for plant growth and development. Members of the HAK\/KUP\/KT (HAK) gene family serve pivotal roles in K+ uptake, translocation and homeostasis. Although numerous HAK genes have been extensively identified across diverse plant species, a comprehensive genomic and functional analysis of this family in sweet potato (Ipomoea batatas L.) remains lacking. In this study, 22 putative IbHAK genes were identified and classified into four distinct clades (I\u2013IV). A systematic characterization was performed for each IbHAK gene, including protein physicochemical properties, chromosome distribution, gene structure, synteny, and promoter cis-elements. Notably, five IbHAK5 genes (IbHAK5A\u2013IbHAK5E) clustered on the HAK gene tree with AtHAK5, OsHAK5, and ZmHAK5. This suggests that small-scale duplication events likely drove the expansion of HAK5 in sweet potato. Among them, IbHAK5A, a gene with broad expression across tissues and strong transcriptional induction under low-K+ (LK) stress, was cloned. The function was then characterized in a K transporter-deficient yeast mutant and an Arabidopsis hak5 mutant. Transcription factor IbPTL1 directly binds the IbHAK5A promoter, upregulates its expression, and integrates into the K+ signaling. In this work, we provide foundational insights into the underlying molecular mechanisms governing K+ acquisition in sweet potato.<\/jats:p>","DOI":"10.3390\/plants15162451","type":"journal-article","created":{"date-parts":[[2026,8,12]],"date-time":"2026-08-12T07:57:26Z","timestamp":1786521446000},"page":"2451","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Identification of the HAK\/KUP\/KT Potassium Transporter Gene Family in Sweet Potato and Functional Characterization of IbHAK5A"],"prefix":"10.3390","volume":"15","author":[{"given":"Fang","family":"Wang","sequence":"first","affiliation":[{"name":"Crop Germplasm Innovation and Genetic Improvement Key Laboratory of Sichuan Province, Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610066, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Zhongmei","family":"Xie","sequence":"additional","affiliation":[{"name":"College of Biological Sciences, China Agricultural University, Beijing 100193, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0002-2874-5509","authenticated-orcid":false,"given":"Songtao","family":"Yang","sequence":"additional","affiliation":[{"name":"Crop Germplasm Innovation and Genetic Improvement Key Laboratory of Sichuan Province, Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610066, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shuai","family":"Qiao","sequence":"additional","affiliation":[{"name":"Crop Germplasm Innovation and Genetic Improvement Key Laboratory of Sichuan Province, Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610066, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Changfeng","family":"Yang","sequence":"additional","affiliation":[{"name":"Crop Germplasm Innovation and Genetic Improvement Key Laboratory of Sichuan Province, Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610066, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Cuiping","family":"Li","sequence":"additional","affiliation":[{"name":"Crop Germplasm Innovation and Genetic Improvement Key Laboratory of Sichuan Province, Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610066, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wei","family":"Song","sequence":"additional","affiliation":[{"name":"Crop Germplasm Innovation and Genetic Improvement Key Laboratory of Sichuan Province, Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610066, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wenfang","family":"Tan","sequence":"additional","affiliation":[{"name":"Crop Germplasm Innovation and Genetic Improvement Key Laboratory of Sichuan Province, Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610066, China"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2026,8,12]]},"reference":[{"key":"ref_1","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1111\/j.1469-8137.1984.tb04103.x","article-title":"A hypothesis relating critical potassium concentrations for growth to the distribution and functions of this ion in the plant cell","volume":"97","author":"Leigh","year":"1984","journal-title":"New Phytol."},{"key":"ref_2","doi-asserted-by":"crossref","first-page":"34","DOI":"10.1111\/jipb.13053","article-title":"Potassium and phosphorus transport and signaling in plants","volume":"63","author":"Wang","year":"2021","journal-title":"J. Integr. Plant Biol."},{"key":"ref_3","doi-asserted-by":"crossref","unstructured":"Ragel, P., Raddatz, N., Leidi, E.O., Quintero, F.J., and Pardo, J.M. (2019). Regulation of K+ nutrition in plants. Front. Plant Sci., 10.","DOI":"10.3389\/fpls.2019.00281"},{"key":"ref_4","doi-asserted-by":"crossref","first-page":"250","DOI":"10.1016\/j.pbi.2009.04.003","article-title":"Physiological functions of mineral macronutrients","volume":"12","author":"Maathuis","year":"2009","journal-title":"Curr. Opin. Plant Biol."},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"863","DOI":"10.1104\/pp.110.154369","article-title":"High-affinity K(+) transport in Arabidopsis: AtHAK5 and AKT1 are vital for seedling establishment and postgermination growth under low-potassium conditions","volume":"153","author":"Pyo","year":"2010","journal-title":"Plant Physiol."},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"10510","DOI":"10.1073\/pnas.93.19.10510","article-title":"Potassium homeostasis in vacuolate plant cells","volume":"93","author":"Walker","year":"1996","journal-title":"Proc. Natl. Acad. Sci. USA"},{"key":"ref_7","doi-asserted-by":"crossref","first-page":"451","DOI":"10.1146\/annurev-arplant-050312-120153","article-title":"Potassium transport and signaling in higher plants","volume":"64","author":"Wang","year":"2013","journal-title":"Annu. Rev. Plant Biol."},{"key":"ref_8","doi-asserted-by":"crossref","first-page":"441","DOI":"10.1146\/annurev.bb.23.060194.002301","article-title":"Perspectives on the physiology and structure of inward-rectifying K+ channels in higher plants: Biophysical implications for K+ uptake","volume":"23","author":"Schroeder","year":"1994","journal-title":"Annu. Rev. Biophys. Biomol. Struct."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"656","DOI":"10.1016\/j.jplph.2013.08.008","article-title":"Potassium in agriculture--status and perspectives","volume":"171","author":"Zorb","year":"2014","journal-title":"Plant Physiol."},{"key":"ref_10","doi-asserted-by":"crossref","first-page":"833","DOI":"10.1093\/jxb\/ert402","article-title":"Molecular mechanisms involved in plant adaptation to low K(+) availability","volume":"65","author":"Cherel","year":"2014","journal-title":"J. Exp. Bot."},{"key":"ref_11","doi-asserted-by":"crossref","first-page":"123","DOI":"10.1016\/j.pbi.2017.06.006","article-title":"Regulation of potassium transport and signaling in plants","volume":"39","author":"Wang","year":"2017","journal-title":"Curr. Opin. Plant Biol."},{"key":"ref_12","doi-asserted-by":"crossref","first-page":"6925","DOI":"10.1128\/jb.175.21.6925-6931.1993","article-title":"Nucleotide sequence and 3\u2032-end deletion studies indicate that the K(+)-uptake protein kup from Escherichia coli is composed of a hydrophobic core linked to a large and partially essential hydrophilic C terminus","volume":"175","author":"Schleyer","year":"1993","journal-title":"J. Bacteriol."},{"key":"ref_13","doi-asserted-by":"crossref","first-page":"3021","DOI":"10.1002\/j.1460-2075.1995.tb07304.x","article-title":"A potassium transporter of the yeast Schwanniomyces occidentalis homologous to the Kup system of Escherichia coli has a high concentrative capacity","volume":"14","author":"Klein","year":"1995","journal-title":"EMBO J."},{"key":"ref_14","first-page":"1","article-title":"Potassium transport in fungi and plants","volume":"1469","year":"2000","journal-title":"Biochim. Biophys."},{"key":"ref_15","first-page":"2281","article-title":"The HAK1 gene of barley is a member of a large gene family and encodes a high-affinity potassium transporter","volume":"9","author":"Rubio","year":"1997","journal-title":"Plant Cell"},{"key":"ref_16","doi-asserted-by":"crossref","first-page":"206","DOI":"10.1016\/S0014-5793(97)01125-3","article-title":"A new family of K+ transporters from Arabidopsis that are conserved across phyla","volume":"415","author":"Quintero","year":"1997","journal-title":"FEBS Lett."},{"key":"ref_17","first-page":"63","article-title":"AtKuP1: A dual-affinity K+ transporter from Arabidopsis","volume":"10","author":"Fu","year":"1998","journal-title":"Plant Cell"},{"key":"ref_18","doi-asserted-by":"crossref","first-page":"945","DOI":"10.1104\/pp.114.246520","article-title":"The role of a potassium transporter OsHAK5 in potassium acquisition and transport from roots to shoots in rice at low potassium supply levels","volume":"166","author":"Yang","year":"2014","journal-title":"Plant Physiol."},{"key":"ref_19","doi-asserted-by":"crossref","first-page":"2747","DOI":"10.1111\/pce.12585","article-title":"Rice potassium transporter OsHAK1 is essential for maintaining potassium-mediated growth and functions in salt tolerance over low and high potassium concentration ranges","volume":"38","author":"Chen","year":"2015","journal-title":"Plant Cell Environ."},{"key":"ref_20","doi-asserted-by":"crossref","first-page":"691","DOI":"10.1111\/jipb.12756","article-title":"ZmHAK5 and ZmHAK1 function in K+ uptake and distribution in maize under low K+ conditions","volume":"61","author":"Qin","year":"2019","journal-title":"J. Integr. Plant Biol."},{"key":"ref_21","doi-asserted-by":"crossref","unstructured":"Cheng, X., Liu, X., Mao, W., Zhang, X., Chen, S., Zhan, K., Bi, H., and Xu, H. (2018). Genome-wide identification and analysis of HAK\/KUP\/KT potassium transporters gene family in wheat (Triticum aestivum L.). Int. J. Mol. Sci., 19.","DOI":"10.3390\/ijms19123969"},{"key":"ref_22","doi-asserted-by":"crossref","unstructured":"Yang, T., Lu, X., Wang, Y., Xie, Y., Ma, J., Cheng, X., Xia, E., Wan, X., and Zhang, Z. (2020). HAK\/KUP\/KT family potassium transporter genes are involved in potassium deficiency and stress responses in tea plants (Camellia sinensis L.): Expression and functional analysis. BMC Genom., 21.","DOI":"10.1186\/s12864-020-06948-6"},{"key":"ref_23","doi-asserted-by":"crossref","unstructured":"Zhou, J., Zhou, H.J., Chen, P., Zhang, L.L., Zhu, J.T., Li, P.-F., Yang, J., Ke, Y.-Z., Zhou, Y.-H., and Li, J.-N. (2020). Genome-wide survey and expression analysis of the KT\/HAK\/KUP family in Brassica napus and its potential roles in the response to K(+) deficiency. Int. J. Mol. Sci., 21.","DOI":"10.3390\/ijms21249487"},{"key":"ref_24","doi-asserted-by":"crossref","unstructured":"Feng, X., Wang, Y., Zhang, N., Wu, Z., Zeng, Q., Wu, J., Wu, X., Wang, L., Zhang, J., and Qi, Y. (2020). Genome-wide systematic characterization of the HAK\/KUP\/KT gene family and its expression profile during plant growth and in response to low-K(+) stress in Saccharum. BMC Plant Biol., 20.","DOI":"10.1186\/s12870-019-2227-7"},{"key":"ref_25","doi-asserted-by":"crossref","first-page":"15","DOI":"10.1270\/jsbbs.16126","article-title":"Improvement for agronomically important traits by gene engineering in sweetpotato","volume":"67","author":"Liu","year":"2017","journal-title":"Breed. Sci."},{"key":"ref_26","doi-asserted-by":"crossref","unstructured":"Lv, Z., and Lu, G. (2021). A new curve of critical leaf potassium concentration based on the maximum root dry matter for diagnosing potassium nutritional status of sweet potato. Front. Plant Sci., 12.","DOI":"10.3389\/fpls.2021.714279"},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"1057","DOI":"10.3724\/SP.J.1006.2017.01057","article-title":"Effect of potassium application on root grow and yield of sweet potato and its physiological mechanism","volume":"43","author":"Wang","year":"2017","journal-title":"Acta Agron. Sin."},{"key":"ref_28","doi-asserted-by":"crossref","unstructured":"Gao, Y., Tang, Z., Xia, H., Sheng, M., Liu, M., Pan, S., Li, Z., and Liu, J. (2021). Potassium fertilization stimulates sucrose-to-starch conversion and root formation in sweet potato (Ipomoea batatas (L.) Lam.). Int. J. Mol. Sci., 22.","DOI":"10.3390\/ijms22094826"},{"key":"ref_29","doi-asserted-by":"crossref","first-page":"77","DOI":"10.1016\/j.jplph.2018.04.008","article-title":"KT-HAK-KUP transporters in major terrestrial photosynthetic organisms: A twenty years tale","volume":"226","author":"Oliferuk","year":"2018","journal-title":"Plant Physiol."},{"key":"ref_30","doi-asserted-by":"crossref","first-page":"2584","DOI":"10.1111\/nph.20319","article-title":"Alternating inverse modulation of xylem K+\/NO3\u2212 loading by HY5 and PIF facilitates diurnal regulation of root-to-shoot water and nutrient transport","volume":"245","author":"Jing","year":"2025","journal-title":"New Phytol."},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"kiaf684","DOI":"10.1093\/plphys\/kiaf684","article-title":"Phosphorylation of GhSKD1 by GhCIPK6D1 regulates potassium efflux during the drought response in cotton","volume":"200","author":"Sun","year":"2026","journal-title":"Plant Physiol."},{"key":"ref_32","doi-asserted-by":"crossref","first-page":"2935","DOI":"10.1111\/pce.70165","article-title":"Potassium Deficiency and Hormone Signalling in Plants","volume":"49","author":"Mishra","year":"2025","journal-title":"Plant Cell Environ."},{"key":"ref_33","doi-asserted-by":"crossref","unstructured":"Wang, F., Tan, W.F., Song, W., Yang, S.T., and Qiao, S. (2022). Transcriptome analysis of sweet potato responses to potassium deficiency. BMC Genom., 23.","DOI":"10.1186\/s12864-022-08870-5"},{"key":"ref_34","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1007\/s13205-020-02552-3","article-title":"Genome-wide characterization and expression analysis of HAK K(+) transport family in Ipomoea","volume":"11","author":"Jin","year":"2021","journal-title":"3 Biotech"},{"key":"ref_35","doi-asserted-by":"crossref","first-page":"25","DOI":"10.1007\/s11248-025-00437-w","article-title":"The high-affinity K+ transporter IbHAK5 enhances potassium ion absorption and improves root morphology in sweetpotato (Ipomoea batatas)","volume":"34","author":"Jin","year":"2025","journal-title":"Transgenic Res."},{"key":"ref_36","doi-asserted-by":"crossref","unstructured":"Cai, K., Zeng, F., Wang, J., and Zhang, G. (2021). Identification and characterization of HAK\/KUP\/KT potassium transporter gene family in barley and their expression under abiotic stress. BMC Genom., 22.","DOI":"10.1186\/s12864-021-07633-y"},{"key":"ref_37","doi-asserted-by":"crossref","unstructured":"Liu, K., Qin, Y., Wu, L., Yi, R., Shi, X., Yu, J., Shi, X., Dong, W., Jia, L., and Fan, M. (2025). Genome-wide identification, characterization and expression pattern analysis of HAK\/KUP\/KT potassium transporter gene family in potato. Front. Plant Sci., 15.","DOI":"10.3389\/fpls.2024.1487794"},{"key":"ref_38","doi-asserted-by":"crossref","unstructured":"Azeem, F., Ijaz, U., Ali, M.A., Hussain, S., Zubair, M., Manzoor, H., Abid, M., Zameer, R., Kim, D.-S., and Golokhvast, K.S. (2021). Genome-wide identification and expression profiling of potassium transport-related genes in Vigna radiata under abiotic stresses. Plants, 11.","DOI":"10.3390\/plants11010002"},{"key":"ref_39","doi-asserted-by":"crossref","unstructured":"Ou, W., Mao, X., Huang, C., Tie, W., Yan, Y., Ding, Z., Wu, C., Xia, Z., Wang, W., and Zhou, S. (2018). Genome-wide identification and expression analysis of the KUP Family under abiotic stress in Cassava (Manihot esculenta Crantz). Front. Physiol., 9.","DOI":"10.3389\/fphys.2018.00017"},{"key":"ref_40","doi-asserted-by":"crossref","first-page":"109642","DOI":"10.1016\/j.plaphy.2025.109642","article-title":"Genome-wide analysis of PHT gene family and their role in LP and salt stress in sweet potato","volume":"221","author":"Wang","year":"2025","journal-title":"Plant Physiol. Biochem."},{"key":"ref_41","first-page":"2863","article-title":"The CBL-interacting protein kinase CIPK23 regulates HAK5-mediated high-affinity K+ uptake in Arabidopsis roots","volume":"169","author":"Ragel","year":"2015","journal-title":"Plant Physiol."},{"key":"ref_42","doi-asserted-by":"crossref","first-page":"5053","DOI":"10.1093\/jxb\/eraa212","article-title":"Arabidopsis K+ transporter HAK5-mediated high-affinity root K+ uptake is regulated by protein kinases CIPK1 and CIPK9","volume":"71","author":"Lara","year":"2020","journal-title":"J. Exp. Bot."},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"1042","DOI":"10.1093\/mp\/sss003","article-title":"The Arabidopsis AP2\/ERF transcription factor RAP2.11 modulates plant response to low-potassium conditions","volume":"5","author":"Kim","year":"2012","journal-title":"Mol. Plant"},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"176","DOI":"10.1111\/nph.17589","article-title":"MYB77 regulates high-affinity potassium uptake by promoting expression of HAK5","volume":"232","author":"Feng","year":"2021","journal-title":"New Phytol."},{"key":"ref_45","doi-asserted-by":"crossref","first-page":"3005","DOI":"10.1105\/tpc.16.00684","article-title":"Phosphorylation of ARF2 relieves its repression of transcription of the K+ transporter gene HAK5 in response to low potassium stress","volume":"28","author":"Zhao","year":"2016","journal-title":"Plant Cell"},{"key":"ref_46","doi-asserted-by":"crossref","unstructured":"Zhao, X., Liu, Y., Liu, X., and Jiang, J. (2018). Comparative transcriptome profiling of two tomato genotypes in response to potassium-deficiency stress. Int. J. Mol. Sci., 19.","DOI":"10.3390\/ijms19082402"},{"key":"ref_47","doi-asserted-by":"crossref","unstructured":"Zhao, Y., Sun, R., Liu, H., Liu, X., Xu, K., Xiao, K., Zhang, S., Yang, X., and Xue, C. (2020). Multi-omics analyses reveal the molecular mechanisms underlying the adaptation of wheat (Triticum aestivum L.) to potassium deprivation. Front. Plant Sci., 11.","DOI":"10.3389\/fpls.2020.588994"},{"key":"ref_48","doi-asserted-by":"crossref","first-page":"3022","DOI":"10.1093\/molbev\/msab120","article-title":"MEGA11: Molecular evolutionary genetics analysis version 11","volume":"38","author":"Tamura","year":"2021","journal-title":"Mol. Biol. Evol."},{"key":"ref_49","doi-asserted-by":"crossref","first-page":"1194","DOI":"10.1016\/j.molp.2020.06.009","article-title":"TBtools: An integrative toolkit developed for interactive analyses of big biological data","volume":"13","author":"Chen","year":"2020","journal-title":"Mol. Plant"},{"key":"ref_50","doi-asserted-by":"crossref","first-page":"e49","DOI":"10.1093\/nar\/gkr1293","article-title":"MCScanX: A toolkit for detection and evolutionary analysis of gene synteny and collinearity","volume":"40","author":"Wang","year":"2012","journal-title":"Nucleic Acids Res."},{"key":"ref_51","first-page":"2848","article-title":"TRK1 encodes a plasma membrane protein required for high-affinity potassium transport in Saccharomyces cerevisiae","volume":"8","author":"Gaber","year":"1988","journal-title":"Mol. Cell Biol."},{"key":"ref_52","doi-asserted-by":"crossref","first-page":"696","DOI":"10.1038\/s41477-017-0002-z","article-title":"Haplotype-resolved sweet potato genome traces back its hexaploidization history","volume":"3","author":"Yang","year":"2017","journal-title":"Nat. Plants"}],"container-title":["Plants"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.mdpi.com\/2223-7747\/15\/16\/2451\/pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,8,14]],"date-time":"2026-08-14T04:21:38Z","timestamp":1786681298000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.mdpi.com\/2223-7747\/15\/16\/2451"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,12]]},"references-count":52,"journal-issue":{"issue":"16","published-online":{"date-parts":[[2026,8]]}},"alternative-id":["plants15162451"],"URL":"https:\/\/doi.org\/10.3390\/plants15162451","relation":{},"ISSN":["2223-7747"],"issn-type":[{"value":"2223-7747","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,8,12]]}}}