{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,5]],"date-time":"2026-10-05T21:37:42Z","timestamp":1791236262887,"version":"4.1.0"},"reference-count":115,"publisher":"MDPI AG","issue":"8","license":[{"start":{"date-parts":[[2014,8,13]],"date-time":"2014-08-13T00:00:00Z","timestamp":1407888000000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/3.0\/"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Viruses"],"abstract":"<jats:p>Bats are being increasingly recognized as reservoir hosts of highly pathogenic and zoonotic emerging viruses (Marburg virus, Nipah virus, Hendra virus, Rabies virus, and coronaviruses). While numerous studies have focused on the mentioned highly  human-pathogenic bat viruses in tropical regions, little is known on similar  human-pathogenic viruses that may be present in European bats. Although novel viruses are being detected, their zoonotic potential remains unclear unless further studies are  conducted. At present, it is assumed that the risk posed by bats to the general public is rather low. In this review, selected viruses detected and isolated in Europe are discussed from our point of view in regard to their human-pathogenic potential. All European bat species and their roosts are legally protected and some European species are even endangered.  Nevertheless, the increasing public fear of bats and their viruses is an obstacle to their  protection. Educating the public regarding bat lyssaviruses might result in reduced threats to both the public and the bats.<\/jats:p>","DOI":"10.3390\/v6083110","type":"journal-article","created":{"date-parts":[[2014,8,13]],"date-time":"2014-08-13T12:15:41Z","timestamp":1407932141000},"page":"3110-3128","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":54,"title":["European Bats as Carriers of Viruses with Zoonotic Potential"],"prefix":"10.3390","volume":"6","author":[{"given":"Claudia","family":"Kohl","sequence":"first","affiliation":[{"name":"Centre for Biological Threats and Special Pathogens, Robert Koch Institute, Nordufer 20,  13353 Berlin, Germany"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Andreas","family":"Kurth","sequence":"additional","affiliation":[{"name":"Centre for Biological Threats and Special Pathogens, Robert Koch Institute, Nordufer 20,  13353 Berlin, Germany"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2014,8,13]]},"reference":[{"key":"ref_1","unstructured":"International Union for Conservation of Nature and Natural Resources (IUCN) 2013 The IUCN Red List of Threatened Species. Version 2013.2. Available online:http:\/\/www.iucnredlist.org\/."},{"key":"ref_2","unstructured":"CMS Convention on the Conservation of Migratory Species of Wild Animals. Available online:http:\/\/www.cms.int\/."},{"key":"ref_3","doi-asserted-by":"crossref","first-page":"531","DOI":"10.1128\/CMR.00017-06","article-title":"Bats: Important reservoir hosts of emerging viruses","volume":"19","author":"Calisher","year":"2006","journal-title":"Clin. Microbiol. Rev."},{"key":"ref_4","first-page":"521","article-title":"Bats prove to be rich reservoirs for emerging viruses","volume":"3","author":"Calisher","year":"2008","journal-title":"Microbe"},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"7159","DOI":"10.3402\/ehtj.v4i0.7159","article-title":"Bats, emerging infectious diseases, and the rabies paradigm revisited","volume":"4","author":"Kuzmin","year":"2011","journal-title":"Emerg. Health Threats J."},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"84","DOI":"10.1016\/j.coviro.2012.11.006","article-title":"Bats and their virome: An important source of emerging viruses capable of infecting humans","volume":"3","author":"Smith","year":"2013","journal-title":"Curr. Opin. Virol."},{"key":"ref_7","unstructured":"Smith, G., and Kelly, A.M. (2008). Food Security in a Global Economy Veterinary Medicine and Public Health, University of Pennsylvania Press. Chapter 13."},{"key":"ref_8","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1007\/s12250-009-3032-5","article-title":"Bats and viruses: A brief review","volume":"24","author":"Wang","year":"2009","journal-title":"Virol. Sin."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"649","DOI":"10.1016\/j.coviro.2011.10.013","article-title":"Mass extinctions, biodiversity and mitochondrial function: Are bats \u201cspecial\u201d as reservoirs for emerging viruses?","volume":"1","author":"Wang","year":"2011","journal-title":"Curr. Opin. Virol."},{"key":"ref_10","doi-asserted-by":"crossref","first-page":"67","DOI":"10.1002\/rmv.520","article-title":"Bats as a continuing source of emerging infections in humans","volume":"17","author":"Wong","year":"2007","journal-title":"Rev. Med. Virol."},{"key":"ref_11","doi-asserted-by":"crossref","unstructured":"Chen, L., Liu, B., Yang, J., and Jin, Q. (2014). DBatVir: The database of bat-associated viruses. Database Oxford, 2014.","DOI":"10.1093\/database\/bau021"},{"key":"ref_12","doi-asserted-by":"crossref","first-page":"2974","DOI":"10.3390\/v6082974","article-title":"Lyssaviruses and Bats: Emergence and Zoonotic Threat","volume":"6","author":"Banyard","year":"2014","journal-title":"Viruses"},{"key":"ref_13","doi-asserted-by":"crossref","first-page":"2052","DOI":"10.3201\/eid1512.090646","article-title":"New adenovirus in bats, Germany","volume":"15","author":"Sonntag","year":"2009","journal-title":"Emerg. Infect. Dis."},{"key":"ref_14","doi-asserted-by":"crossref","first-page":"1888","DOI":"10.1128\/JVI.05974-11","article-title":"Genome analysis of bat adenovirus 2: Indications of interspecies transmission","volume":"86","author":"Kohl","year":"2012","journal-title":"J. Virol."},{"key":"ref_15","doi-asserted-by":"crossref","first-page":"118","DOI":"10.1016\/j.tvjl.2010.06.020","article-title":"Novel adenoviruses and herpesviruses detected in bats","volume":"189","author":"Vidovszky","year":"2011","journal-title":"Vet. J."},{"key":"ref_16","doi-asserted-by":"crossref","first-page":"449","DOI":"10.3201\/eid1703.100526","article-title":"Amplification of emerging viruses in a bat colony","volume":"17","author":"Drexler","year":"2011","journal-title":"Emerg. Infect. Dis."},{"key":"ref_17","doi-asserted-by":"crossref","first-page":"95","DOI":"10.4149\/av_2014_01_95","article-title":"Novel European lineages of bat astroviruses identified in Hungary","volume":"58","author":"Kemenesi","year":"2014","journal-title":"Acta Virol."},{"key":"ref_18","doi-asserted-by":"crossref","first-page":"e87194","DOI":"10.1371\/journal.pone.0087194","article-title":"A preliminary study of viral metagenomics of French bat species in contact with humans: Identification of new mammalian viruses","volume":"9","author":"Dacheux","year":"2014","journal-title":"PLoS One"},{"key":"ref_19","doi-asserted-by":"crossref","first-page":"433","DOI":"10.4269\/ajtmh.1988.38.433","article-title":"Ecology of viruses isolated from sand flies in Italy and characterized of a new Phlebovirus (Arabia virus)","volume":"38","author":"Verani","year":"1988","journal-title":"Am. J. Trop. Med. Hyg."},{"key":"ref_20","doi-asserted-by":"crossref","first-page":"626","DOI":"10.3201\/eid1404.071439","article-title":"Detection and prevalence patterns of group I coronaviruses in bats, northern Germany","volume":"14","author":"Ipsen","year":"2008","journal-title":"Emerg. Infect. Dis."},{"key":"ref_21","doi-asserted-by":"crossref","first-page":"1883","DOI":"10.1007\/s00705-011-1057-1","article-title":"Detection of alpha and betacoronaviruses in multiple Iberian bat species","volume":"156","author":"Casas","year":"2011","journal-title":"Arch. Virol."},{"key":"ref_22","doi-asserted-by":"crossref","first-page":"2679","DOI":"10.3390\/v5112679","article-title":"Detection of coronaviruses in bats of various species in Italy","volume":"5","author":"Lelli","year":"2013","journal-title":"Viruses"},{"key":"ref_23","doi-asserted-by":"crossref","first-page":"530","DOI":"10.1089\/vbz.2011.0829","article-title":"Alphacoronavirus detected in bats in the United Kingdom","volume":"12","author":"August","year":"2012","journal-title":"Vector Borne Zoonotic Dis."},{"key":"ref_24","doi-asserted-by":"crossref","first-page":"11336","DOI":"10.1128\/JVI.00650-10","article-title":"Genomic characterization of severe acute respiratory syndrome-related coronavirus in European bats and classification of coronaviruses based on partial RNA-dependent RNA polymerase gene sequences","volume":"84","author":"Drexler","year":"2010","journal-title":"J. Virol."},{"key":"ref_25","doi-asserted-by":"crossref","first-page":"507","DOI":"10.1007\/s00705-010-0612-5","article-title":"Identification of SARS-like coronaviruses in horseshoe bats (Rhinolophus hipposideros) in Slovenia","volume":"155","author":"Rihtaric","year":"2010","journal-title":"Arch. Virol."},{"key":"ref_26","doi-asserted-by":"crossref","first-page":"456","DOI":"10.3201\/eid1903.121503","article-title":"Human betacoronavirus 2c EMC\/2012-related viruses in bats, Ghana and Europe","volume":"19","author":"Annan","year":"2013","journal-title":"Emerg. Infect. Dis."},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"785","DOI":"10.1089\/vbz.2009.0173","article-title":"Circulation of group 2 coronaviruses in a bat species common to urban areas in Western Europe","volume":"10","author":"Reusken","year":"2010","journal-title":"Vector Borne Zoonotic Dis."},{"key":"ref_28","doi-asserted-by":"crossref","first-page":"366","DOI":"10.1007\/s11262-013-1008-x","article-title":"Alpha and lineage C betaCoV infections in Italian bats","volume":"48","author":"Marciano","year":"2014","journal-title":"Virus Genes"},{"key":"ref_29","doi-asserted-by":"crossref","first-page":"e1002304","DOI":"10.1371\/journal.ppat.1002304","article-title":"Discovery of an Ebolavirus-like filovirus in Europe","volume":"7","author":"Negredo","year":"2011","journal-title":"PLoS Pathog."},{"key":"ref_30","doi-asserted-by":"crossref","first-page":"9134","DOI":"10.1128\/JVI.00800-12","article-title":"Bats worldwide carry hepatitis E-related viruses that form a putative novel genus within the family Hepeviridae","volume":"86","author":"Drexler","year":"2012","journal-title":"J. Virol."},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"2651","DOI":"10.1099\/vir.0.83045-0","article-title":"Discovery of herpesviruses in bats","volume":"88","author":"Wibbelt","year":"2007","journal-title":"J. Gen. Virol."},{"key":"ref_32","doi-asserted-by":"crossref","first-page":"529","DOI":"10.1556\/avet.56.2008.4.10","article-title":"Detection of a novel bat gammaherpesvirus in Hungary","volume":"56","author":"Harrach","year":"2008","journal-title":"Acta Vet. Hung."},{"key":"ref_33","doi-asserted-by":"crossref","first-page":"94","DOI":"10.1093\/gbe\/evt211","article-title":"Novel papillomaviruses in free-ranging Iberian bats: No virus-host co-evolution, no strict host specificity, and hints for recombination","volume":"6","author":"Garin","year":"2014","journal-title":"Genome Biol. Evol."},{"key":"ref_34","doi-asserted-by":"crossref","first-page":"e38688","DOI":"10.1371\/journal.pone.0038688","article-title":"Novel paramyxoviruses in free-ranging European bats","volume":"7","author":"Kurth","year":"2012","journal-title":"PLoS One"},{"key":"ref_35","doi-asserted-by":"crossref","first-page":"796","DOI":"10.1038\/ncomms1796","article-title":"Bats host major mammalian paramyxoviruses","volume":"3","author":"Drexler","year":"2012","journal-title":"Nat. Commun."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"e43106","DOI":"10.1371\/journal.pone.0043106","article-title":"Isolation and characterization of three mammalian orthoreoviruses from European bats","volume":"7","author":"Kohl","year":"2012","journal-title":"PLoS One"},{"key":"ref_37","doi-asserted-by":"crossref","first-page":"84","DOI":"10.1111\/zph.12001","article-title":"Identification of Mammalian Orthoreovirus Type 3 in Italian Bats","volume":"60","author":"Lelli","year":"2012","journal-title":"Zoonoses Public Health"},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"2419","DOI":"10.1128\/jcm.30.9.2419-2426.1992","article-title":"Antigenic and molecular characterization of bat rabies virus in Europe","volume":"30","author":"Bourhy","year":"1992","journal-title":"J. Clin. Microbiol."},{"key":"ref_39","doi-asserted-by":"crossref","first-page":"1029","DOI":"10.1017\/S0950268803001481","article-title":"European bat lyssaviruses: An emerging zoonosis","volume":"131","author":"Fooks","year":"2003","journal-title":"Epidemiol. Infect."},{"key":"ref_40","doi-asserted-by":"crossref","first-page":"22","DOI":"10.1111\/zph.12002","article-title":"Bat rabies surveillance in Europe","volume":"60","author":"Schatz","year":"2013","journal-title":"Zoonoses Public Health"},{"key":"ref_41","doi-asserted-by":"crossref","first-page":"1854","DOI":"10.3201\/eid1112.041200","article-title":"European bat lyssaviruses, The Netherlands","volume":"11","author":"Verstraten","year":"2005","journal-title":"Emerg. Infect. Dis."},{"key":"ref_42","doi-asserted-by":"crossref","first-page":"e2057","DOI":"10.1371\/journal.pone.0002057","article-title":"Genomic diversity and evolution of the lyssaviruses","volume":"3","author":"Delmas","year":"2008","journal-title":"PLoS One"},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"9557","DOI":"10.1128\/JVI.00771-10","article-title":"Application of broad-spectrum resequencing microarray for genotyping rhabdoviruses","volume":"84","author":"Dacheux","year":"2010","journal-title":"J. Virol."},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"3268","DOI":"10.1128\/JVI.75.7.3268-3276.2001","article-title":"Evidence of two Lyssavirus phylogroups with distinct pathogenicity and immunogenicity","volume":"75","author":"Badrane","year":"2001","journal-title":"J. Virol."},{"key":"ref_45","doi-asserted-by":"crossref","first-page":"273","DOI":"10.1007\/s00705-006-0853-5","article-title":"Epidemiology of bat rabies in Germany","volume":"152","author":"Johnson","year":"2007","journal-title":"Arch. Virol."},{"key":"ref_46","doi-asserted-by":"crossref","first-page":"589","DOI":"10.1136\/vr.154.19.589","article-title":"Genetic analysis of European bat lyssavirus type 1 isolates from France","volume":"154","author":"Barrat","year":"2004","journal-title":"Vet. Rec."},{"key":"ref_47","doi-asserted-by":"crossref","first-page":"2319","DOI":"10.1099\/0022-1317-78-9-2319","article-title":"Evolution of European bat lyssaviruses","volume":"78","author":"Amengual","year":"1997","journal-title":"J. Gen. Virol."},{"key":"ref_48","first-page":"1519","article-title":"Novel lyssavirus in Natterer\u2019s bat, Germany","volume":"17","author":"Freuling","year":"2011","journal-title":"Emerg. Infect. Dis."},{"key":"ref_49","doi-asserted-by":"crossref","first-page":"1581","DOI":"10.1017\/S0950268810000373","article-title":"First encounter of European bat lyssavirus type 2 (EBLV-2) in a bat in Finland","volume":"138","author":"Lilley","year":"2010","journal-title":"Epidemiol. Infect."},{"key":"ref_50","doi-asserted-by":"crossref","first-page":"520","DOI":"10.3201\/eid1703.100894","article-title":"Phylogeny of European bat Lyssavirus 1 in Eptesicus isabellinus bats, Spain","volume":"17","author":"Juste","year":"2011","journal-title":"Emerg. Infect. Dis."},{"key":"ref_51","doi-asserted-by":"crossref","first-page":"383","DOI":"10.1136\/vr.152.13.383","article-title":"Isolation of a European bat lyssavirus type 2 from a Daubenton\u2019s bat in the United Kingdom","volume":"152","author":"Johnson","year":"2003","journal-title":"Vet. Rec."},{"key":"ref_52","doi-asserted-by":"crossref","first-page":"2333","DOI":"10.1007\/s00705-013-1747-y","article-title":"Isolation of Bokeloh bat lyssavirus in Myotis nattereri in France","volume":"158","author":"Servat","year":"2013","journal-title":"Arch. Virol."},{"key":"ref_53","doi-asserted-by":"crossref","first-page":"201","DOI":"10.1016\/j.virusres.2013.07.021","article-title":"Molecular diagnostics for the detection of Bokeloh bat lyssavirus in a bat from Bavaria, Germany","volume":"177","author":"Freuling","year":"2013","journal-title":"Virus Res."},{"key":"ref_54","doi-asserted-by":"crossref","first-page":"69","DOI":"10.1099\/vir.0.046490-0","article-title":"Detection of rhabdovirus viral RNA in oropharyngeal swabs and ectoparasites of Spanish bats","volume":"94","author":"Marston","year":"2013","journal-title":"J. Gen. Virol."},{"key":"ref_55","doi-asserted-by":"crossref","first-page":"490","DOI":"10.1007\/BF01765224","article-title":"Die aetiologie der tuberkulose","volume":"11","author":"Koch","year":"1932","journal-title":"Klin. Wochenschr."},{"key":"ref_56","unstructured":"International Committee on Taxonomy of Viruses (ICTV) online 2013. Available online:http:\/\/www.ictvonline.org\/virusTaxonomy.asp?version=2013."},{"key":"ref_57","doi-asserted-by":"crossref","first-page":"1967","DOI":"10.1056\/NEJMoa030747","article-title":"Identification of a novel coronavirus in patients with severe acute respiratory syndrome","volume":"348","author":"Drosten","year":"2003","journal-title":"N. Engl. J. Med."},{"key":"ref_58","doi-asserted-by":"crossref","first-page":"240","DOI":"10.1038\/423240a","article-title":"Aetiology: Koch\u2019s postulates fulfilled for SARS virus","volume":"423","author":"Fouchier","year":"2003","journal-title":"Nature"},{"key":"ref_59","doi-asserted-by":"crossref","first-page":"263","DOI":"10.1016\/S0140-6736(03)13967-0","article-title":"Newly discovered coronavirus as the primary cause of severe acute respiratory syndrome","volume":"362","author":"Kuiken","year":"2003","journal-title":"Lancet"},{"key":"ref_60","doi-asserted-by":"crossref","first-page":"S88","DOI":"10.1038\/nm1143","article-title":"Severe acute respiratory syndrome","volume":"10","author":"Peiris","year":"2004","journal-title":"Nat. Med."},{"key":"ref_61","doi-asserted-by":"crossref","first-page":"1834","DOI":"10.3201\/eid1212.060401","article-title":"Review of bats and SARS","volume":"12","author":"Wang","year":"2006","journal-title":"Emerg. Infect. Dis."},{"key":"ref_62","unstructured":"WHO. Available online:http:\/\/www.who.int\/csr\/don\/2014_06_26_mers\/en."},{"key":"ref_63","doi-asserted-by":"crossref","unstructured":"The WHO MERS-CoV Research Group (2013). State of knowledge and data gaps of Middle East respiratory syndrome coronavirus (MERS-CoV) in humans. PLOS Curr., 5, doi:10.1371\/currents.outbreaks.0bf719e352e7478f8ad85fa30127ddb8.","DOI":"10.1371\/currents.outbreaks.0bf719e352e7478f8ad85fa30127ddb8"},{"key":"ref_64","doi-asserted-by":"crossref","first-page":"e01146\u201314","DOI":"10.1128\/mBio.01146-14","article-title":"Middle East respiratory syndrome coronavirus quasispecies that include homologues of human isolates revealed through whole-genome analysis and virus cultured from dromedary camels in Saudi Arabia","volume":"5","author":"Briese","year":"2014","journal-title":"MBio"},{"key":"ref_65","doi-asserted-by":"crossref","first-page":"1697","DOI":"10.3201\/eid1910.130946","article-title":"Close relative of human Middle East respiratory syndrome coronavirus in bat, South Africa","volume":"19","author":"Ithete","year":"2013","journal-title":"Emerg. Infect. Dis."},{"key":"ref_66","doi-asserted-by":"crossref","first-page":"4211","DOI":"10.1128\/JVI.80.9.4211-4219.2006","article-title":"Animal origins of the severe acute respiratory syndrome coronavirus: Insight from ACE2-S-protein interactions","volume":"80","author":"Li","year":"2006","journal-title":"J. Virol."},{"key":"ref_67","doi-asserted-by":"crossref","first-page":"535","DOI":"10.1038\/nature12711","article-title":"Isolation and characterization of a bat SARS-like coronavirus that uses the ACE2 receptor","volume":"503","author":"Ge","year":"2013","journal-title":"Nature"},{"key":"ref_68","doi-asserted-by":"crossref","first-page":"7","DOI":"10.3201\/eid1911.131172","article-title":"Middle East respiratory syndrome coronavirus in bats, Saudi Arabia","volume":"19","author":"Memish","year":"2013","journal-title":"Emerg. Infect. Dis."},{"key":"ref_69","doi-asserted-by":"crossref","unstructured":"Mole, B. (2013). Deadly coronavirus found in bats. Nat. News.","DOI":"10.1038\/nature.2013.13597"},{"key":"ref_70","doi-asserted-by":"crossref","first-page":"2025","DOI":"10.1098\/rstb.2010.0037","article-title":"Predicting species distribution and abundance responses to climate change: Why it is essential to include biotic interactions across trophic levels","volume":"365","author":"Macel","year":"2010","journal-title":"Philos. Trans. R. Soc. Lond. B Biol. Sci."},{"key":"ref_71","unstructured":"(1999). ArcGIS Explorer, ESRI Inc."},{"key":"ref_72","doi-asserted-by":"crossref","first-page":"431","DOI":"10.1016\/j.ympev.2003.08.017","article-title":"Molecular systematics and biogeography of the bent-wing bat complex Miniopterus schreibersii (Kuhl, 1817) (Chiroptera: Vespertilionidae)","volume":"31","author":"Appleton","year":"2004","journal-title":"Mol. Phylogenet. Evol."},{"key":"ref_73","doi-asserted-by":"crossref","first-page":"e29773","DOI":"10.1371\/journal.pone.0029773","article-title":"Diseases and causes of death in European bats: Dynamics in disease susceptibility and infection rates","volume":"6","author":"Speck","year":"2011","journal-title":"PLoS One"},{"key":"ref_74","doi-asserted-by":"crossref","unstructured":"Fereidouni, S., Kwasnitschka, L., Balkema Buschmann, A., M\u00fcller, T., Freuling, C., Schatz, J., Pikula, J., Bandouchova, H., Hoffmann, R., and Ohlendorf, B. (2014). No Virological Evidence for an Influenza A\u2014Like Virus in European Bats. Zoonoses Public Health.","DOI":"10.1111\/zph.12131"},{"key":"ref_75","doi-asserted-by":"crossref","first-page":"16151","DOI":"10.1073\/pnas.1308049110","article-title":"Bats carry pathogenic hepadnaviruses antigenically related to hepatitis B virus and capable of infecting human hepatocytes","volume":"110","author":"Drexler","year":"2013","journal-title":"Proc. Natl. Acad. Sci. USA"},{"key":"ref_76","unstructured":"Kohl, C., and Kurth, A. (2013). Unpublished work."},{"key":"ref_77","first-page":"231","article-title":"Genomic characterization of M and S RNA segments of hantaviruses isolated from bats","volume":"39","author":"Jung","year":"1995","journal-title":"Acta Virol."},{"key":"ref_78","unstructured":"Kohl, C., Vidovszky, M.Z., and Kurth, A. Available online:http:\/\/talk.ictvonline.org\/."},{"key":"ref_79","doi-asserted-by":"crossref","first-page":"355","DOI":"10.1051\/vetres:2006058","article-title":"Canine respiratory viruses","volume":"38","author":"Buonavoglia","year":"2007","journal-title":"Vet. Res."},{"key":"ref_80","doi-asserted-by":"crossref","first-page":"464","DOI":"10.1016\/j.meegid.2006.02.007","article-title":"The evolutionary history and dynamics of bat rabies virus","volume":"6","author":"Davis","year":"2006","journal-title":"Infect. Genet. Evol."},{"key":"ref_81","doi-asserted-by":"crossref","first-page":"19","DOI":"10.1016\/j.vetmic.2005.05.014","article-title":"Virological and molecular characterization of a mammalian orthoreovirus type 3 strain isolated from a dog in Italy","volume":"109","author":"Decaro","year":"2005","journal-title":"Vet. Microbiol."},{"key":"ref_82","doi-asserted-by":"crossref","first-page":"1664","DOI":"10.1086\/383129","article-title":"Isolation and molecular characterization of a novel type 3 reovirus from a child with meningitis","volume":"189","author":"Tyler","year":"2004","journal-title":"J. Infect. Dis."},{"key":"ref_83","doi-asserted-by":"crossref","first-page":"3818","DOI":"10.1128\/JCM.01531-13","article-title":"High similarity of novel orthoreovirus detected in a child hospitalized with acute gastroenteritis to mammalian orthoreoviruses found in bats in Europe","volume":"51","author":"Steyer","year":"2013","journal-title":"J. Clin. Microbiol."},{"key":"ref_84","doi-asserted-by":"crossref","first-page":"11424","DOI":"10.1073\/pnas.0701372104","article-title":"A previously unknown reovirus of bat origin is associated with an acute respiratory disease in humans","volume":"104","author":"Chua","year":"2007","journal-title":"Proc. Natl. Acad. Sci. USA"},{"key":"ref_85","doi-asserted-by":"crossref","first-page":"e3803","DOI":"10.1371\/journal.pone.0003803","article-title":"Identification and characterization of a new orthoreovirus from patients with acute respiratory infections","volume":"3","author":"Chua","year":"2008","journal-title":"PLoS One"},{"key":"ref_86","doi-asserted-by":"crossref","first-page":"151","DOI":"10.1016\/j.vetmic.2010.02.001","article-title":"Human rabies due to lyssavirus infection of bat origin","volume":"142","author":"Johnson","year":"2010","journal-title":"Vet. Microbiol."},{"key":"ref_87","doi-asserted-by":"crossref","first-page":"58","DOI":"10.1111\/j.1863-2378.2012.01533.x","article-title":"Bat rabies, public health and European bat conservation","volume":"60","author":"Racey","year":"2013","journal-title":"Zoonoses Public Health"},{"key":"ref_88","doi-asserted-by":"crossref","first-page":"378","DOI":"10.1016\/S0140-6736(86)92336-6","article-title":"Human rabies of bat origin in Europe","volume":"1","author":"Lumio","year":"1986","journal-title":"Lancet"},{"key":"ref_89","doi-asserted-by":"crossref","first-page":"281","DOI":"10.1002\/jmv.10481","article-title":"Case report: Isolation of a European bat lyssavirus type 2a from a fatal human case of rabies encephalitis","volume":"71","author":"Fooks","year":"2003","journal-title":"J. Med. Virol."},{"key":"ref_90","unstructured":"Dietz, C., von Helversen, O., and Nill, D. (2007). Handbuch der Flederm\u00e4use Europas und Nordwestafrikas, Kosmos. (in German)."},{"key":"ref_91","doi-asserted-by":"crossref","first-page":"413","DOI":"10.3201\/eid0804.010263","article-title":"European bat lyssavirus infection in Spanish bat populations","volume":"8","author":"Amengual","year":"2002","journal-title":"Emerg. Infect. Dis."},{"key":"ref_92","doi-asserted-by":"crossref","first-page":"49","DOI":"10.1111\/j.1439-0450.2003.00725.x","article-title":"Spill-over of European bat lyssavirus type 1 into a stone marten (Martes foina) in Germany","volume":"51","author":"Cox","year":"2004","journal-title":"J. Vet. Med. B"},{"key":"ref_93","doi-asserted-by":"crossref","first-page":"190","DOI":"10.1016\/j.jcpa.2005.10.005","article-title":"Natural and experimental infection of sheep with European bat lyssavirus type-1 of Danish bat origin","volume":"134","author":"Tjornehoj","year":"2006","journal-title":"J. Comp. Pathol."},{"key":"ref_94","doi-asserted-by":"crossref","first-page":"280","DOI":"10.3201\/eid1502.080637","article-title":"European bat Lyssavirus transmission among cats, Europe","volume":"15","author":"Dacheux","year":"2009","journal-title":"Emerg. Infect. Dis."},{"key":"ref_95","first-page":"3384","article-title":"Ecological and anthropogenic drivers of rabies exposure in vampire bats: Implications for transmission and control","volume":"279","author":"Streicker","year":"2012","journal-title":"Proc. Biol. Sci."},{"key":"ref_96","unstructured":"Cottasche, J.G. (1826). Ansichten der Natur, mit wissenschaftlichen Erl\u00e4uterungen, Buchhandlung. [2nd ed.]. (in German)."},{"key":"ref_97","first-page":"2131","article-title":"Phylogeny, niche conservatism and the latitudinal diversity gradient in mammals","volume":"277","author":"Buckley","year":"2010","journal-title":"Proc. Biol. Sci."},{"key":"ref_98","unstructured":"Wilson, D.E., and Reeder, D.M. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference, Johns Hopkins University. [3rd ed.]."},{"key":"ref_99","doi-asserted-by":"crossref","first-page":"580","DOI":"10.1126\/science.1105113","article-title":"A molecular phylogeny for bats illuminates biogeography and the fossil record","volume":"307","author":"Teeling","year":"2005","journal-title":"Science"},{"key":"ref_100","doi-asserted-by":"crossref","first-page":"189","DOI":"10.2307\/2845093","article-title":"Bat species density gradients in the New World: A statistical assessment","volume":"16","author":"Willig","year":"1989","journal-title":"J. Biogeogr."},{"key":"ref_101","doi-asserted-by":"crossref","first-page":"825","DOI":"10.1046\/j.1461-0248.2003.00504.x","article-title":"Reverse latitudinal trends in species richness of pitcher-plant food webs","volume":"6","author":"Buckley","year":"2003","journal-title":"Ecol. Lett."},{"key":"ref_102","doi-asserted-by":"crossref","first-page":"626","DOI":"10.1073\/pnas.0507535103","article-title":"The diversity and biogeography of soil bacterial communities","volume":"103","author":"Fierer","year":"2006","journal-title":"Proc. Natl. Acad. Sci. USA"},{"key":"ref_103","doi-asserted-by":"crossref","first-page":"251","DOI":"10.1034\/j.1600-0587.2001.240302.x","article-title":"Body size determines the strength of the latitudinal diversity gradient","volume":"24","author":"Hillebrand","year":"2001","journal-title":"Ecography"},{"key":"ref_104","doi-asserted-by":"crossref","first-page":"102","DOI":"10.1038\/nrmicro1341","article-title":"Microbial biogeography: Putting microorganisms on the map","volume":"4","author":"Martiny","year":"2006","journal-title":"Nat. Rev. Microbiol."},{"key":"ref_105","doi-asserted-by":"crossref","first-page":"483","DOI":"10.1007\/s00114-008-0352-5","article-title":"Biogeography of diseases: A framework for analysis","volume":"95","author":"Peterson","year":"2008","journal-title":"Naturwissenschaften"},{"key":"ref_106","doi-asserted-by":"crossref","first-page":"278","DOI":"10.1016\/j.tim.2005.04.003","article-title":"Here a virus, there a virus, everywhere the same virus?","volume":"13","author":"Breitbart","year":"2005","journal-title":"Trends Microbiol."},{"key":"ref_107","doi-asserted-by":"crossref","first-page":"199","DOI":"10.1111\/j.1365-2699.2005.01406.x","article-title":"Microbial biogeography?","volume":"33","author":"Dolan","year":"2006","journal-title":"J. Biogeogr."},{"key":"ref_108","doi-asserted-by":"crossref","first-page":"647","DOI":"10.1038\/nature09575","article-title":"Impacts of biodiversity on the emergence and transmission of infectious diseases","volume":"468","author":"Keesing","year":"2010","journal-title":"Nature"},{"key":"ref_109","doi-asserted-by":"crossref","first-page":"9","DOI":"10.1080\/14888386.2006.9712789","article-title":"Biodiversity loss and emerging infectious disease: An example from the rodent-borne hemorrhagic fevers","volume":"7","author":"Mills","year":"2006","journal-title":"Biodiversity"},{"key":"ref_110","doi-asserted-by":"crossref","first-page":"40","DOI":"10.1111\/j.1469-0691.2008.02691.x","article-title":"Biodiversity loss and the rise of zoonotic pathogens","volume":"15","author":"Ostfeld","year":"2009","journal-title":"Clin. Microbiol. Infect."},{"key":"ref_111","doi-asserted-by":"crossref","first-page":"847","DOI":"10.1017\/S0031182012000200","article-title":"Pangloss revisited: A critique of the dilution effect and the biodiversity-buffers-disease paradigm","volume":"139","author":"Randolph","year":"2012","journal-title":"Parasitology"},{"key":"ref_112","doi-asserted-by":"crossref","first-page":"679","DOI":"10.1111\/ele.12101","article-title":"A meta-analysis suggesting that the relationship between biodiversity and risk of zoonotic pathogen transmission is idiosyncratic","volume":"16","author":"Salkeld","year":"2013","journal-title":"Ecol. Lett."},{"key":"ref_113","doi-asserted-by":"crossref","first-page":"1842","DOI":"10.3201\/eid1112.050997","article-title":"Host range and emerging and reemerging pathogens","volume":"11","author":"Woolhouse","year":"2005","journal-title":"Emerg. Infect. Dis."},{"key":"ref_114","doi-asserted-by":"crossref","first-page":"1109","DOI":"10.1126\/science.1059026","article-title":"Population biology of multihost pathogens","volume":"292","author":"Woolhouse","year":"2001","journal-title":"Science"},{"key":"ref_115","unstructured":"Baas Becking, L.G.M. (1934). Geobiologie of Inleiding tot de Milieukunde, W.P. Van Stockum & Zoon. (in Dutch)."}],"container-title":["Viruses"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.mdpi.com\/1999-4915\/6\/8\/3110\/pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,10,11]],"date-time":"2025-10-11T21:14:44Z","timestamp":1760217284000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.mdpi.com\/1999-4915\/6\/8\/3110"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2014,8,13]]},"references-count":115,"journal-issue":{"issue":"8","published-online":{"date-parts":[[2014,8]]}},"alternative-id":["v6083110"],"URL":"https:\/\/doi.org\/10.3390\/v6083110","relation":{},"ISSN":["1999-4915"],"issn-type":[{"value":"1999-4915","type":"electronic"}],"subject":[],"published":{"date-parts":[[2014,8,13]]}}}