{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,4,23]],"date-time":"2026-04-23T14:04:07Z","timestamp":1776953047971,"version":"3.51.4"},"reference-count":75,"publisher":"MDPI AG","issue":"5","license":[{"start":{"date-parts":[[2024,5,7]],"date-time":"2024-05-07T00:00:00Z","timestamp":1715040000000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"name":"General Secretariat for Research and Technology of Greece","award":["PRIMA2019-11"],"award-info":[{"award-number":["PRIMA2019-11"]}]},{"name":"General Secretariat for Research and Technology of Greece","award":["PRIMA\/0009\/2019"],"award-info":[{"award-number":["PRIMA\/0009\/2019"]}]},{"DOI":"10.13039\/100007340","name":"PRIMA foundation in FCT Portugal","doi-asserted-by":"publisher","award":["PRIMA2019-11"],"award-info":[{"award-number":["PRIMA2019-11"]}],"id":[{"id":"10.13039\/100007340","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100007340","name":"PRIMA foundation in FCT Portugal","doi-asserted-by":"publisher","award":["PRIMA\/0009\/2019"],"award-info":[{"award-number":["PRIMA\/0009\/2019"]}],"id":[{"id":"10.13039\/100007340","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Horticulturae"],"abstract":"<jats:p>In the current study, the effects of drought stress on the growth and phytochemical profile of Scolymus hispanicus L. (a.k.a. golden thistle) were evaluated. Plants were treated with three irrigation regimes, e.g., plants that received only rainwater (Control; C), deficit irrigation (I1; 50% of field capacity (FC)), and full irrigation (\u03992; 100% of FC). The fresh weight of the rosette of leaves was not negatively impacted by deficit irrigation, whereas root development was severely restrained compared to control and I2 treatments. Drought stress conditions had a positive effect on the nutritional properties of the golden thistle since the treatments of control and deficit irrigation showed the highest content of macronutrients and energy. Oxalic acid was the richest organic acid, especially under the I1 regime. Similarly, \u03b1-tocopherol was the only identified vitamin E isoform, whose content was also doubled in I1 treatment. Raffinose, glucose, and sucrose were the most abundant free sugars in amounts that varied among the irrigation treatments, while the total and distinct free sugar content was the highest for the I1 treatment. The most abundant detected fatty acid compounds were \u03b1-linolenic acid, followed by palmitic and linoleic acid, with the highest amount being detected in C, I1, and I2 treatments, respectively. Flavonoids were the only class of polyphenols detected in golden thistle leaves, including mostly kaempferol and quercetin derivatives. The greatest antioxidant potency was shown for the control and I1 treatments (for OxHLIA and TBARS methods, respectively). The evaluated leaf samples recorded a varied antimicrobial effect for the different bacterial strains and fungi, whereas no cytotoxic, hepatotoxic, and anti-inflammatory effects against the tested cell lines were recorded. Finally, the mineral content of leaves was significantly affected by the irrigation regime, with Ca, Mg, Cu, and Zn being the highest for the I1 treatment, while the I2 treatment had the highest content of K, Fe, and Mn and the lowest Na content. In conclusion, deficit irrigation showed promising results since it improved the phytochemical content without compromising the fresh weight of leaves, and thus it could be suggested as a sustainable agronomic practice for producing high-added value products without significant constraints in growth development and yield parameters of golden thistle.<\/jats:p>","DOI":"10.3390\/horticulturae10050479","type":"journal-article","created":{"date-parts":[[2024,5,7]],"date-time":"2024-05-07T09:03:38Z","timestamp":1715072618000},"page":"479","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":7,"title":["The Impact of Deficit Irrigation on the Agronomic Performance and Chemical Composition of Scolymus hispanicus L."],"prefix":"10.3390","volume":"10","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-4535-3075","authenticated-orcid":false,"given":"Nikolaos","family":"Polyzos","sequence":"first","affiliation":[{"name":"Laboratory of Vegetable Production, Department of Agriculture, Crop Production and Rural Environment, University of Thessaly, Fytokou Street, 384 46 Volos, Greece"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5813-4453","authenticated-orcid":false,"given":"Beatriz H.","family":"Paschoalinotto","sequence":"additional","affiliation":[{"name":"Centro de Investiga\u00e7\u00e3o de Montanha (CIMO), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"},{"name":"Laborat\u00f3rio Associado para a Sustentabilidade e Tecnologia em Regi\u00f5es de Montanha (SusTEC), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-3954-3833","authenticated-orcid":false,"given":"T\u00e2nia C. S. P.","family":"Pires","sequence":"additional","affiliation":[{"name":"Centro de Investiga\u00e7\u00e3o de Montanha (CIMO), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"},{"name":"Laborat\u00f3rio Associado para a Sustentabilidade e Tecnologia em Regi\u00f5es de Montanha (SusTEC), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2835-5079","authenticated-orcid":false,"given":"Mikel","family":"A\u00f1ibarro-Ortega","sequence":"additional","affiliation":[{"name":"Centro de Investiga\u00e7\u00e3o de Montanha (CIMO), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"},{"name":"Laborat\u00f3rio Associado para a Sustentabilidade e Tecnologia em Regi\u00f5es de Montanha (SusTEC), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-6801-4578","authenticated-orcid":false,"given":"Ricardo","family":"Calhelha","sequence":"additional","affiliation":[{"name":"Centro de Investiga\u00e7\u00e3o de Montanha (CIMO), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"},{"name":"Laborat\u00f3rio Associado para a Sustentabilidade e Tecnologia em Regi\u00f5es de Montanha (SusTEC), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-4910-4882","authenticated-orcid":false,"given":"Isabel C. F. R.","family":"Ferreira","sequence":"additional","affiliation":[{"name":"Centro de Investiga\u00e7\u00e3o de Montanha (CIMO), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8744-7814","authenticated-orcid":false,"given":"Maria In\u00eas","family":"Dias","sequence":"additional","affiliation":[{"name":"Centro de Investiga\u00e7\u00e3o de Montanha (CIMO), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"},{"name":"Laborat\u00f3rio Associado para a Sustentabilidade e Tecnologia em Regi\u00f5es de Montanha (SusTEC), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9050-5189","authenticated-orcid":false,"given":"Lillian","family":"Barros","sequence":"additional","affiliation":[{"name":"Centro de Investiga\u00e7\u00e3o de Montanha (CIMO), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"},{"name":"Laborat\u00f3rio Associado para a Sustentabilidade e Tecnologia em Regi\u00f5es de Montanha (SusTEC), Instituto Polit\u00e9cnico de Bragan\u00e7a, Campus de Santa Apol\u00f3nia, 5300-253 Bragan\u00e7a, Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-0324-7960","authenticated-orcid":false,"given":"Spyridon A.","family":"Petropoulos","sequence":"additional","affiliation":[{"name":"Laboratory of Vegetable Production, Department of Agriculture, Crop Production and Rural Environment, University of Thessaly, Fytokou Street, 384 46 Volos, Greece"}]}],"member":"1968","published-online":{"date-parts":[[2024,5,7]]},"reference":[{"key":"ref_1","doi-asserted-by":"crossref","first-page":"4","DOI":"10.1007\/s13280-016-0793-6","article-title":"Sustainable intensification of agriculture for human prosperity and global sustainability","volume":"46","author":"Williams","year":"2017","journal-title":"Ambio"},{"key":"ref_2","doi-asserted-by":"crossref","unstructured":"Kumar, V., Kumar, P., and Khan, A. (2020). Optimization of PGPR and silicon fertilization using response surface methodology for enhanced growth, yield and biochemical parameters of French bean (Phaseolus vulgaris L.) under saline stress. Biocatal. Agric. Biotechnol., 23.","DOI":"10.1016\/j.bcab.2019.101463"},{"key":"ref_3","first-page":"100","article-title":"Climate change impact on crop rotations of winter durum wheat and tomato in Southern Italy: Yield analysis and soil fertility","volume":"7","author":"Ventrella","year":"2012","journal-title":"Ital. J. Agron."},{"key":"ref_4","doi-asserted-by":"crossref","first-page":"311","DOI":"10.1016\/j.agwat.2017.08.015","article-title":"Deficit irrigation and emerging fruit crops as a strategy to save water in Mediterranean semiarid agrosystems","volume":"202","author":"Galindo","year":"2018","journal-title":"Agric. Water Manag."},{"key":"ref_5","first-page":"44","article-title":"Future climate change in the Mediterranean area: Implications for water use and weed management","volume":"7","author":"Lovelli","year":"2012","journal-title":"Ital. J. Agron."},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"680","DOI":"10.1017\/S0021859618000126","article-title":"Improvement of plant performance under water deficit with the employment of biological and chemical priming agents","volume":"156","author":"Balestrini","year":"2018","journal-title":"J. Agric. Sci."},{"key":"ref_7","doi-asserted-by":"crossref","first-page":"17","DOI":"10.3389\/fsufs.2020.00017","article-title":"Weak and Strong Sustainability of Irrigation: A Framework for Irrigation Practices Under Limited Water Availability","volume":"4","author":"Borsato","year":"2020","journal-title":"Front. Sustain. Food Syst."},{"key":"ref_8","doi-asserted-by":"crossref","first-page":"119940","DOI":"10.1016\/j.jclepro.2019.119940","article-title":"Modelling crop yield, water consumption, and water use efficiency for sustainable agroecosystem management","volume":"253","author":"Liu","year":"2020","journal-title":"J. Clean. Prod."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"102170","DOI":"10.1016\/j.gloenvcha.2020.102170","article-title":"Mapping global patterns of land use decision-making","volume":"65","author":"Malek","year":"2020","journal-title":"Glob. Environ. Chang."},{"key":"ref_10","doi-asserted-by":"crossref","first-page":"528","DOI":"10.1016\/j.agwat.2009.03.023","article-title":"Improving agricultural water productivity: Between optimism and caution","volume":"97","author":"Molden","year":"2010","journal-title":"Agric. Water Manag."},{"key":"ref_11","doi-asserted-by":"crossref","first-page":"975","DOI":"10.3390\/w7030975","article-title":"Water scarcity and future challenges for food production","volume":"7","author":"Mancosu","year":"2015","journal-title":"Water"},{"key":"ref_12","doi-asserted-by":"crossref","first-page":"1421","DOI":"10.1111\/j.1672-9072.2007.00556.x","article-title":"Deficit irrigation as a strategy to save water: Physiology and potential application to horticulture","volume":"49","author":"Costa","year":"2007","journal-title":"J. Integr. Plant Biol."},{"key":"ref_13","doi-asserted-by":"crossref","unstructured":"Ouda, S., Zohry, A.E.H., and Noreldin, T. (2020). Deficit Irrigation: A Remedy for Water Scarcity, Springer.","DOI":"10.1007\/978-3-030-35586-9"},{"key":"ref_14","doi-asserted-by":"crossref","first-page":"147","DOI":"10.1093\/jxb\/erl165","article-title":"Deficit irrigation for reducing agricultural water use","volume":"58","author":"Fereres","year":"2007","journal-title":"J. Exp. Bot."},{"key":"ref_15","doi-asserted-by":"crossref","first-page":"72","DOI":"10.1016\/j.postharvbio.2011.02.005","article-title":"Activity of extracts from wild edible herbs against postharvest fungal diseases of fruit and vegetables","volume":"61","author":"Gatto","year":"2011","journal-title":"Postharvest Biol. Technol."},{"key":"ref_16","doi-asserted-by":"crossref","first-page":"43","DOI":"10.1016\/j.agwat.2005.02.004","article-title":"Trickle and sprinkler irrigation of potato (Solanum tuberosum L.) in the Middle Anatolian Region in Turkey","volume":"79","author":"Kanber","year":"2006","journal-title":"Agric. Water Manag."},{"key":"ref_17","unstructured":"Goodwin, I., and Boland, A.-M. (2002). Deficit Irrigation Practices, Food and Agriculture Organization of the United Nations (FAO)."},{"key":"ref_18","doi-asserted-by":"crossref","unstructured":"Karkanis, A., Polyzos, N., Kompocholi, M., and Petropoulos, S.A. (2022). Rock samphire, a candidate crop for saline agriculture: Cropping practices, chemical composition and health effects. Appl. Sci., 12.","DOI":"10.3390\/app12020737"},{"key":"ref_19","doi-asserted-by":"crossref","unstructured":"Paschoalinotto, B.H., Polyzos, N., Compocholi, M., Rouphael, Y., Alexopoulos, A., Dias, M.I., Barros, L., and Petropoulos, S.A. (2023). Domestication of Wild Edible Species: The Response of Scolymus hispanicus Plants to Different Fertigation Regimes. Horticulturae, 9.","DOI":"10.3390\/horticulturae9010103"},{"key":"ref_20","doi-asserted-by":"crossref","unstructured":"Petropoulos, S.A., Fernandes, \u00c2., Dias, M.I., Pereira, C., Calhelha, R., Di Gioia, F., Tzortzakis, N., Ivanov, M., Sokovic, M., and Barros, L. (2020). Wild and cultivated Centaurea raphanina subsp. mixta: A valuable source of bioactive compounds. Antioxidants, 9.","DOI":"10.3390\/antiox9040314"},{"key":"ref_21","doi-asserted-by":"crossref","unstructured":"Polyzos, N., Paschoalinotto, B.H., Compocholi, M., Pinela, J., Heleno, S.A., Calhelha, R.C., Dias, M.I., Barros, L., and Petropoulos, S.A. (2022). Fertilization of pot-grown Cichorium spinosum L.: How it can affect plant growth, chemical profile, and bioactivities of edible parts?. Horticulturae, 8.","DOI":"10.3390\/horticulturae8100890"},{"key":"ref_22","unstructured":"Preedy, V., and Watson, R. (2020). The Mediterranean Diet: An Evidence-Based Approach, Academic Press."},{"key":"ref_23","doi-asserted-by":"crossref","unstructured":"Carrascosa, \u00c1., Pascual, A., Ros, M., Petropoulos, S., and Alguacil, M. (2022). The Effect of Fertilization Regime on Growth Parameters of Sonchus oleraceus and Two Genotypes of Portulaca oleracea. Biol. Life Sci. Forum, 16.","DOI":"10.3390\/IECHo2022-12515"},{"key":"ref_24","doi-asserted-by":"crossref","first-page":"69","DOI":"10.1016\/j.tifs.2018.02.006","article-title":"Edible halophytes of the Mediterranean basin: Potential candidates for novel food products","volume":"74","author":"Petropoulos","year":"2018","journal-title":"Trends Food Sci. Technol."},{"key":"ref_25","doi-asserted-by":"crossref","unstructured":"Renna, M. (2018). Reviewing the prospects of sea fennel (Crithmum maritimum L.) as emerging vegetable crop. Plants, 7.","DOI":"10.3390\/plants7040092"},{"key":"ref_26","doi-asserted-by":"crossref","unstructured":"Zenobi, S., Fiorentini, M., Ledda, L., Deligios, P., Aquilanti, L., and Orsini, R. (2022). Crithmum maritimum L. Biomass Production in Mediterranean Environment. Agronomy, 12.","DOI":"10.3390\/agronomy12040926"},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"111182","DOI":"10.1016\/j.scienta.2022.111182","article-title":"Effect of moderate salinity on Golden Thistle (Scolymus hispanicus L.) grown in a soilless cropping system","volume":"303","author":"Papadimitriou","year":"2022","journal-title":"Sci. Hortic."},{"key":"ref_28","doi-asserted-by":"crossref","first-page":"2669","DOI":"10.1080\/14786419.2017.1283494","article-title":"Phytochemical and pharmacological variability in golden thistle functional parts: Comparative study of roots, stems, leaves and flowers","volume":"31","author":"Marmouzi","year":"2017","journal-title":"Nat. Prod. Res."},{"key":"ref_29","doi-asserted-by":"crossref","unstructured":"Lentini, F., and Venza, F. (2007). Wild food plants of popular use in Sicily. J. Ethnobiol. Ethnomed., 3.","DOI":"10.1186\/1746-4269-3-15"},{"key":"ref_30","doi-asserted-by":"crossref","first-page":"130","DOI":"10.1663\/0013-0001(2006)60[130:WGFPIT]2.0.CO;2","article-title":"Wild gathered food plants in the European Mediterranean: A comparative analysis","volume":"60","author":"Leonti","year":"2006","journal-title":"Econ. Bot."},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1111\/j.1095-8339.2006.00549.x","article-title":"Ethnobotanical review of wild edible plants in Spain","volume":"152","author":"Morales","year":"2006","journal-title":"Bot. J. Linn. Soc."},{"key":"ref_32","first-page":"383","article-title":"Qualitative characterisation of cultivated and wild edible plants: Mineral elements, phenols content and antioxidant capacity","volume":"12","author":"Disciglio","year":"2017","journal-title":"Ital. J. Agron."},{"key":"ref_33","doi-asserted-by":"crossref","first-page":"859","DOI":"10.1016\/j.foodres.2018.10.069","article-title":"Bioactive compounds content and antimicrobial activities of wild edible Asteraceae species of the Mediterranean flora under commercial cultivation conditions","volume":"119","author":"Petropoulos","year":"2019","journal-title":"Food Res. Int."},{"key":"ref_34","doi-asserted-by":"crossref","first-page":"431","DOI":"10.1007\/s10722-011-9693-6","article-title":"Wild vegetables of the Mediterranean area as valuable sources of bioactive compounds","volume":"59","author":"Morales","year":"2012","journal-title":"Genet. Resour. Crop Evol."},{"key":"ref_35","doi-asserted-by":"crossref","first-page":"822","DOI":"10.1016\/j.foodchem.2005.08.058","article-title":"Lipid concentrations of wild edible greens in Crete","volume":"99","author":"Vardavas","year":"2006","journal-title":"Food Chem."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"467","DOI":"10.1002\/ptr.1243","article-title":"In vitro antioxidant activity of non-cultivated vegetables of ethnic Albanians in southern Italy","volume":"16","author":"Pieroni","year":"2002","journal-title":"Phytother. Res."},{"key":"ref_37","doi-asserted-by":"crossref","unstructured":"Polo, S., Tard\u00edo, J., V\u00e9lez-del-Burgo, A., Molina, M., and Pardo-de-Santayana, M. (2009). Knowledge, use and ecology of golden thistle (Scolymus hispanicus L.) in Central Spain. J. Ethnobiol. Ethnomed., 5.","DOI":"10.1186\/1746-4269-5-42"},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"281","DOI":"10.1177\/1082013211427798","article-title":"Fatty acids profiles of some Spanish wild vegetables","volume":"18","author":"Morales","year":"2012","journal-title":"Food Sci. Technol. Int."},{"key":"ref_39","doi-asserted-by":"crossref","first-page":"389","DOI":"10.1016\/j.lwt.2013.08.017","article-title":"Mediterranean non-cultivated vegetables as dietary sources of compounds with antioxidant and biological activity","volume":"55","author":"Morales","year":"2014","journal-title":"LWT Food Sci. Technol."},{"key":"ref_40","doi-asserted-by":"crossref","unstructured":"Chaski, C., and Petropoulos, S.A. (2022). The Effects of Biostimulant Application on Growth Parameters of Lettuce Plants Grown under Deficit Irrigation Conditions. Biol. Life Sci. Forum, 16.","DOI":"10.3390\/IECHo2022-12499"},{"key":"ref_41","unstructured":"Horwitz, W., and Latimer, G. (2019). Official Methods of Analysis of Association of Official Analytical Chemists, AOAC Inter."},{"key":"ref_42","doi-asserted-by":"crossref","first-page":"1732","DOI":"10.1016\/j.jff.2013.07.019","article-title":"Bioactivity and chemical characterization in hydrophilic and lipophilic compounds of Chenopodium ambrosioides L.","volume":"5","author":"Barros","year":"2013","journal-title":"J. Funct. Foods"},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"44","DOI":"10.2993\/0278-0771-34.1.44","article-title":"Weeds and food diversity: Natural yield assessment and future alternatives for traditionally consumed wild vegetables","volume":"34","author":"Molina","year":"2014","journal-title":"J. Ethnobiol."},{"key":"ref_44","doi-asserted-by":"crossref","unstructured":"Calone, R., Mircea, D.M., Gonz\u00e1lez-Orenga, S., Boscaiu, M., Lambertini, C., Barbanti, L., and Vicente, O. (2022). Recovery from Salinity and Drought Stress in the Perennial Sarcocornia fruticosa vs. the Annual Salicornia europaea and S. veneta. Plants, 11.","DOI":"10.3390\/plants11081058"},{"key":"ref_45","doi-asserted-by":"crossref","first-page":"163","DOI":"10.1016\/j.jfca.2014.02.009","article-title":"Nutrient composition of six wild edible Mediterranean Asteraceae plants of dietary interest","volume":"34","author":"Molina","year":"2014","journal-title":"J. Food Compos. Anal."},{"key":"ref_46","doi-asserted-by":"crossref","first-page":"129","DOI":"10.1016\/j.foodchem.2016.07.080","article-title":"Salinity effect on nutritional value, chemical composition and bioactive compounds content of Cichorium spinosum L.","volume":"214","author":"Petropoulos","year":"2017","journal-title":"Food Chem."},{"key":"ref_47","doi-asserted-by":"crossref","first-page":"83","DOI":"10.1016\/j.foodchem.2017.05.092","article-title":"Successive harvesting affects yield, chemical composition and antioxidant activity of Cichorium spinosum L.","volume":"237","author":"Petropoulos","year":"2017","journal-title":"Food Chem."},{"key":"ref_48","doi-asserted-by":"crossref","first-page":"813","DOI":"10.1016\/j.foodchem.2005.08.057","article-title":"The antioxidant and phylloquinone content of wildly grown greens in Crete","volume":"99","author":"Vardavas","year":"2006","journal-title":"Food Chem."},{"key":"ref_49","doi-asserted-by":"crossref","first-page":"777","DOI":"10.1111\/ppl.13033","article-title":"Short versus prolonged freezing differentially impacts freeze\u2013thaw injury in spinach leaves: Mechanistic insights through metabolite profiling","volume":"168","author":"Min","year":"2019","journal-title":"Physiol. Plant."},{"key":"ref_50","doi-asserted-by":"crossref","first-page":"223","DOI":"10.21273\/JASHS.135.3.223","article-title":"Regulated Water Deficits Improve Phytochemical Concentration in Lettuce","volume":"135","author":"Oh","year":"2010","journal-title":"J. Am. Soc. Hortic. Sci."},{"key":"ref_51","first-page":"812","article-title":"Nutritional Yield of African Indigenous Vegetables in Water-Deficient and Water-Sufficient Conditions","volume":"05","author":"Luoh","year":"2014","journal-title":"Food Nutr. Sci."},{"key":"ref_52","doi-asserted-by":"crossref","first-page":"263","DOI":"10.4067\/S0716-97602004000200013","article-title":"Omega-3 fatty acids and antioxidants in edible wild plants","volume":"37","author":"Simopoulos","year":"2004","journal-title":"Biol. Res."},{"key":"ref_53","doi-asserted-by":"crossref","first-page":"967","DOI":"10.1007\/s10529-008-9639-z","article-title":"Fatty acid unsaturation, mobilization, and regulation in the response of plants to stress","volume":"30","author":"Upchurch","year":"2008","journal-title":"Biotechnol. Lett."},{"key":"ref_54","doi-asserted-by":"crossref","first-page":"273","DOI":"10.2135\/cropsci2010.06.0368","article-title":"Membrane fatty acid composition and saturation levels associated with leaf dehydration tolerance and post-drought rehydration in Kentucky bluegrass","volume":"51","author":"Xu","year":"2011","journal-title":"Crop Sci."},{"key":"ref_55","doi-asserted-by":"crossref","first-page":"271","DOI":"10.1016\/j.scienta.2008.10.016","article-title":"Water deficit effects on Salvia officinalis fatty acids and essential oils composition","volume":"120","author":"Bettaieb","year":"2009","journal-title":"Sci. Hortic."},{"key":"ref_56","first-page":"414","article-title":"Macroelements and Microelements Content of Some Wild Edible Plants","volume":"24","author":"Ghanimi","year":"2022","journal-title":"Asian J. Microbiol. Biotechnol. Environ. Sci."},{"key":"ref_57","doi-asserted-by":"crossref","first-page":"753","DOI":"10.1007\/s00217-018-3197-y","article-title":"Effect of in vitro gastrointestinal digestion on the total phenolic contents and antioxidant activity of wild Mediterranean edible plant extracts","volume":"245","author":"Moreira","year":"2019","journal-title":"Eur. Food Res. Technol."},{"key":"ref_58","doi-asserted-by":"crossref","first-page":"125","DOI":"10.1016\/j.indcrop.2014.10.021","article-title":"Phenolic profile and antioxidant properties of commercial and wild Fragaria vesca L. roots: A comparison between hydromethanolic and aqueous extracts","volume":"63","author":"Dias","year":"2015","journal-title":"Ind. Crops Prod."},{"key":"ref_59","doi-asserted-by":"crossref","unstructured":"Rolim, P.M., Fidelis, G.P., Padilha, C.E.A., Santos, E.S., Rocha, H.A.O., and Macedo, G.R. (2018). Phenolic profile and antioxidant activity from peels and seeds of melon (Cucumis melo L. var. reticulatus) and their antiproliferative effect in cancer cells. Braz. J. Med. Biol. Res., 51.","DOI":"10.1590\/1414-431x20176069"},{"key":"ref_60","doi-asserted-by":"crossref","first-page":"241","DOI":"10.1016\/j.plaphy.2005.01.020","article-title":"Effect of water and temperature stress on the content of active constituents of Hypericum brasiliense Choisy","volume":"43","author":"Mazzafera","year":"2005","journal-title":"Plant Physiol. Biochem."},{"key":"ref_61","first-page":"217037","article-title":"Reactive Oxygen Species, Oxidative Damage, and Antioxidative Defense Mechanism in Plants under Stressful Conditions","volume":"2012","author":"Sharma","year":"2012","journal-title":"J. Bot."},{"key":"ref_62","doi-asserted-by":"crossref","unstructured":"Hasanuzzaman, M., Bhuyan, M.H.M.B., Zulfiqar, F., Raza, A., Mohsin, S.M., Al Mahmud, J., Fujita, M., and Fotopoulos, V. (2020). Reactive oxygen species and antioxidant defense in plants under abiotic stress: Revisiting the crucial role of a universal defense regulator. Antioxidants, 9.","DOI":"10.3390\/antiox9080681"},{"key":"ref_63","doi-asserted-by":"crossref","first-page":"303","DOI":"10.1016\/j.foodchem.2010.06.037","article-title":"Effect of meteorological conditions on antioxidant flavonoids in Portuguese cultivars of white and red onions","volume":"124","author":"Rodrigues","year":"2011","journal-title":"Food Chem."},{"key":"ref_64","doi-asserted-by":"crossref","unstructured":"Guarise, M., Borgonovo, G., Bassoli, A., and Ferrante, A. (2023). The Effect of Drought on Sisymbrium officinale (L.) Wild Species for Potential Cultivation as a Leafy Vegetable. Horticulturae, 9.","DOI":"10.3390\/horticulturae9010111"},{"key":"ref_65","doi-asserted-by":"crossref","first-page":"108078","DOI":"10.1016\/j.agwat.2022.108078","article-title":"Effect of water regime, nitrogen level and biostimulants application on yield and quality traits of wild rocket [Diplotaxis tenuifolia (L.) DC.]","volume":"277","author":"Schiattone","year":"2023","journal-title":"Agric. Water Manag."},{"key":"ref_66","doi-asserted-by":"crossref","first-page":"7974","DOI":"10.1002\/fsn3.3716","article-title":"Golden thistle (Scolymus hispanicus L.) hydromethanolic extracts ameliorated glucose absorption and inflammatory markers in vitro","volume":"11","author":"Gulec","year":"2023","journal-title":"Food Sci. Nutr."},{"key":"ref_67","doi-asserted-by":"crossref","first-page":"1103","DOI":"10.1007\/s11738-010-0638-z","article-title":"Drought effects on polyphenol composition and antioxidant activities in aerial parts of Salvia officinalis L.","volume":"33","author":"Bettaieb","year":"2011","journal-title":"Acta Physiol. Plant."},{"key":"ref_68","doi-asserted-by":"crossref","unstructured":"Sergio, L., Boari, F., Pieralice, M., Linsalata, V., Cantore, V., and Venere, D. (2020). Di Bioactive phenolics and antioxidant capacity of some wild edible greens as affected by different cooking treatments. Foods, 9.","DOI":"10.3390\/foods9091320"},{"key":"ref_69","doi-asserted-by":"crossref","first-page":"664","DOI":"10.1016\/j.foodchem.2008.06.027","article-title":"Antioxidant capacity of medicinal plants from the Province of C\u00f3rdoba (Argentina) and their in vitro testing in a model food system","volume":"112","author":"Borneo","year":"2009","journal-title":"Food Chem."},{"key":"ref_70","doi-asserted-by":"crossref","first-page":"999","DOI":"10.1007\/s12010-018-2784-5","article-title":"Drought Stress Effects on Growth, ROS Markers, Compatible Solutes, Phenolics, Flavonoids, and Antioxidant Activity in Amaranthus tricolor","volume":"186","author":"Sarker","year":"2018","journal-title":"Appl. Biochem. Biotechnol."},{"key":"ref_71","doi-asserted-by":"crossref","first-page":"400","DOI":"10.1016\/j.plaphy.2017.04.012","article-title":"Effect of in vitro drought stress on phenolic acids, flavonols, saponins, and antioxidant activity in Agave salmiana","volume":"115","year":"2017","journal-title":"Plant Physiol. Biochem."},{"key":"ref_72","doi-asserted-by":"crossref","first-page":"265","DOI":"10.31989\/ffhd.v10i6.718","article-title":"Screening of the antimicrobial activity of some extracts of edible wild plants in Morocco","volume":"10","author":"Aboukhalaf","year":"2020","journal-title":"Funct. Foods Health Dis."},{"key":"ref_73","doi-asserted-by":"crossref","first-page":"523","DOI":"10.1089\/jmf.2021.0144","article-title":"Effects of Golden Thistle (Scolymus hispanicus L.) on Cytotoxic Activity: Cell Cycle Arrest and Apoptotic Properties on the CaCo-2 Cell Line","volume":"25","author":"Gulec","year":"2022","journal-title":"J. Med. Food"},{"key":"ref_74","doi-asserted-by":"crossref","unstructured":"Petropoulos, S.A., Fernandes, \u00c2., Dias, M.I., Pereira, C., Calhelha, R.C., Ivanov, M., Sokovic, M.D., Ferreira, I.C.F.R., and Barros, L. (2021). Effects of growing substrate and nitrogen fertilization on the chemical composition and bioactive properties of Centaurea raphanina ssp. mixta (DC.) Runemark. Agronomy, 11.","DOI":"10.3390\/agronomy11030576"},{"key":"ref_75","doi-asserted-by":"crossref","unstructured":"Mikropoulou, E.V., Vougogiannopoulou, K., Kalpoutzakis, E., Sklirou, A.D., Skaperda, Z., Houriet, J., Wolfender, J.L., Trougakos, I.P., Kouretas, D., and Halabalaki, M. (2018). Phytochemical composition of the decoctions of Greek edible greens (ch\u00f3rta) and evaluation of antioxidant and cytotoxic properties. 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