{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,8]],"date-time":"2026-09-08T23:31:55Z","timestamp":1788910315364,"version":"build-2803163510"},"reference-count":45,"publisher":"Pensoft Publishers","license":[{"start":{"date-parts":[[2026,6,10]],"date-time":"2026-06-10T00:00:00Z","timestamp":1781049600000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["PK"],"abstract":"<jats:p>\n                    <jats:italic>Timmia<\/jats:italic>\n                    is a small moss genus with only eight currently accepted species. However, the boundaries and taxonomic status of some taxa remain uncertain and controversial, which has significantly hindered studies on its species diversity. During our 2025 expeditions to the Sanjiangyuan Region in Qinghai Province, China, we discovered a moss resembling\n                    <jats:italic>T. sphaerocarpa<\/jats:italic>\n                    with globose capsules, distinguishable by several morphological characters: stomata confined to the capsule base; vegetative and perichaetial leaves with a distinctly differentiated sheathing base; limb laminal cells with distinct mamillae; naked antheridia; sinuose exothecial cells; and spores 2\u20133 times larger than those of\n                    <jats:italic>T. sphaerocarpa<\/jats:italic>\n                    . Morphological and molecular phylogenetic analyses, based on two chloroplast markers (\n                    <jats:italic>atp<\/jats:italic>\n                    B-\n                    <jats:italic>rbc<\/jats:italic>\n                    L spacer and\n                    <jats:italic>trn<\/jats:italic>\n                    L-\n                    <jats:italic>trn<\/jats:italic>\n                    F) and the nuclear 26S region, suggest that this moss represents a new species, herein described as\n                    <jats:italic>T. sanjiangyuanensis<\/jats:italic>\n                    . A new section, sect.\n                    <jats:italic>Pseudosphaerocarpa<\/jats:italic>\n                    , is established to accommodate this species. Descriptions, illustrations and SEM photographs of the peristome and spores of this new species, along with an updated key to\n                    <jats:italic>Timmia<\/jats:italic>\n                    species in China, are provided.\n                  <\/jats:p>","DOI":"10.3897\/phytokeys.276.176402","type":"journal-article","created":{"date-parts":[[2026,6,10]],"date-time":"2026-06-10T13:35:04Z","timestamp":1781098504000},"page":"35-49","source":"Crossref","is-referenced-by-count":1,"title":["Timmia sanjiangyuanensis (Timmiaceae, Bryophyta), a new species from the Qinghai Plateau, China"],"prefix":"10.3897","volume":"276","author":[{"given":"Xin-Yin","family":"Ma","sequence":"first","affiliation":[{"name":"College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou, China"},{"name":"Zhejiang Provincial Key Laboratory of Wetland Intelligent Monitoring and Ecological Restoration, Hangzhou Normal University, Hangzhou, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jia-Yi","family":"Zheng","sequence":"additional","affiliation":[{"name":"College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou, China"},{"name":"Zhejiang Provincial Key Laboratory of Wetland Intelligent Monitoring and Ecological Restoration, Hangzhou Normal University, Hangzhou, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xin-Rui","family":"Xia","sequence":"additional","affiliation":[{"name":"College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou, China"},{"name":"Zhejiang Provincial Key Laboratory of Wetland Intelligent Monitoring and Ecological Restoration, Hangzhou Normal University, Hangzhou, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xiao-Tong","family":"Song","sequence":"additional","affiliation":[{"name":"School of Biotechnology, Jiangnan University, Wuxi, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ya-Kun","family":"Shi","sequence":"additional","affiliation":[{"name":"Northwest Survey and Planning Institute of the National Forestry and Grassland Administration, Xi\u2019an, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yu-Huan","family":"Wu","sequence":"additional","affiliation":[{"name":"College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou, China"},{"name":"Zhejiang Provincial Key Laboratory of Wetland Intelligent Monitoring and Ecological Restoration, Hangzhou Normal University, Hangzhou, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wen-Zhuan","family":"Huang","sequence":"additional","affiliation":[{"name":"College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou, China"},{"name":"Zhejiang Provincial Key Laboratory of Wetland Intelligent Monitoring and Ecological Restoration, Hangzhou Normal University, Hangzhou, China"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"2258","published-online":{"date-parts":[[2026,6,10]]},"reference":[{"key":"176402_B1","doi-asserted-by":"publisher","DOI":"10.31111\/nsnr\/2024.58.2.B39"},{"key":"176402_B2","unstructured":"Brassard GR (1979) The moss genus Timmia. 1. Introduction, and revision of T. norvegica and allied taxa. Lindbergia 5: 39\u201353. https:\/\/www.jstor.org\/stable\/20149323"},{"key":"176402_B3","unstructured":"Brassard GR (1980) The moss genus Timmia. 2. Sect. Timmiaurea. Lindbergia 6: 129\u2013136. https:\/\/www.jstor.org\/stable\/20149368"},{"key":"176402_B4","unstructured":"Brassard GR (1984) The moss genus Timmia. 3. Sect. Timmia. Lindbergia 10: 33\u201344. https:\/\/www.jstor.org\/stable\/20149505"},{"key":"176402_B5","unstructured":"Brassard GR (2007) Timmiaceae Schimper. In: Flora of North America Editorial Committee (Eds) Flora of North America, Vol. 27, part 1. Oxford University Press, Oxford, 165\u2013169."},{"key":"176402_B6","unstructured":"Brinda C, Atwood JJ (2025) The Bryophyte Nomenclator. https:\/\/www.bryonames.org [accessed on 24 Dec. 2025]"},{"key":"176402_B7","unstructured":"Brotherus VF (1929) Musci. 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Borntraeger Verlagsbuchhandlung, Stuttgart, 1\u2013263."},{"key":"176402_B15","doi-asserted-by":"publisher","DOI":"10.1639\/0007-2745-114.4.749"},{"key":"176402_B16","doi-asserted-by":"publisher","DOI":"10.1080\/03736687.2021.1963914"},{"key":"176402_B17","doi-asserted-by":"publisher","DOI":"10.1639\/0007-2745-122.4.597"},{"key":"176402_B18","doi-asserted-by":"publisher","DOI":"10.3390\/plants13131759"},{"key":"176402_B19","doi-asserted-by":"publisher","DOI":"10.1179\/174328206X152289"},{"key":"176402_B20","doi-asserted-by":"publisher","DOI":"10.1186\/s13059-020-02154-5"},{"key":"176402_B21","doi-asserted-by":"publisher","DOI":"10.1038\/nmeth.4285"},{"key":"176402_B22","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/mst010"},{"key":"176402_B23","doi-asserted-by":"publisher","DOI":"10.1093\/bioinformatics\/bts199"},{"key":"176402_B24","unstructured":"Li XJ (2006) Flora Bryophytorum Sinicorum. Vol. 4 Eubryales. Science Press, Beijing, 1\u2013263."},{"key":"176402_B25","unstructured":"Li XJ, He S (2007) Timmiaceae. In: Li XJ, Crosby MR, He S (Eds) Moss Flora of China Vol. 4. Bryaceae\u2013Timmiaceae. Science Press, Beijing & Missouri Botanical Garden, St. Louis, 194\u2013198."},{"key":"176402_B26","doi-asserted-by":"publisher","DOI":"10.7606\/j.issn.1000-4025.2022.12.2152"},{"key":"176402_B27","doi-asserted-by":"publisher","DOI":"10.1179\/jbr.1993.17.3.481"},{"key":"176402_B28","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msaa015"},{"key":"176402_B29","doi-asserted-by":"publisher","DOI":"10.5962\/bhl.title.156377"},{"key":"176402_B30","unstructured":"Mittermeier RA, Robles Gil P, Hoffman M, Pilgrim J, Brooks T, Goettsch Mittermeier C, Lamoreux J, da Fonseca GAB (2004) Hotspots Revisited: Earth\u2019s Biologically Richest and Most Endangered Terrestrial Ecoregions. Conservation International, Mexico, 1\u2013392."},{"key":"176402_B31","unstructured":"M\u00fcller C (1898) Bryologia provinciae Schen-si Sinensis ex collectione Giraldiana III. 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