{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,15]],"date-time":"2026-07-15T19:17:52Z","timestamp":1784143072833,"version":"3.55.0"},"reference-count":41,"publisher":"Pensoft Publishers","license":[{"start":{"date-parts":[[2023,12,20]],"date-time":"2023-12-20T00:00:00Z","timestamp":1703030400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["ZK"],"abstract":"<jats:p><jats:italic>Calotes wangi<\/jats:italic><jats:bold>sp. nov.<\/jats:bold>, a new species of the agamid genus <jats:italic>Calotes<\/jats:italic> Cuvier, 1817, from southern China and northern Vietnam, is described. This species can be distinguished from all known congeners by a combination of morphological characteristics and genetic divergence in the mitochondrial tRNA, ND2, and CO1 genes. Molecular phylogenetic analysis revealed that the new species was formed as a monophyletic group and that considerable genetic divergence existed between its congeners (minimum <jats:italic>p<\/jats:italic>-distance, 4.6%). <jats:italic>Calotes wangi<\/jats:italic><jats:bold>sp. nov.<\/jats:bold> is distinguished by a combination of the following characteristics: average SVL &amp;lt; 90 mm for adult males; 10\u201314 dorsal eyelid scales; scales on side of neck and adjacent shoulder area pointing obliquely upward; keels on neck scales weakly to strongly developed; fold in front of the shoulder absent; pair of dark triangular patches extending from the front of the shoulder to the jaw angles; and orange coloration of the tongue. <jats:italic>Calotes wangi<\/jats:italic><jats:bold>sp. nov.<\/jats:bold> is similar to <jats:italic>C. irawadi<\/jats:italic> but differs in having scales between the nasal shield and the orbit and a fourth toe with a claw that can reach between the eyes and tympanum (even to the snout when hind the limbs are adpressed forward). Phylogenetic analyses revealed two well-supported subspecies, Lineages A and B in <jats:italic>C. wangi<\/jats:italic><jats:bold>sp. nov.<\/jats:bold>, with mean uncorrected <jats:italic>p<\/jats:italic>-distances between them of 2%. We propose that Lineage A, which is mainly from the central and southern Wuzhi Mountains on Hainan Island, is a subspecies, <jats:italic>C. w. hainanensis<\/jats:italic><jats:bold>ssp. nov.<\/jats:bold> Lineage B mainly comprises individuals from other sites on the island plus the adjacent mainland, and is described as subspecies, <jats:italic>C. w. wangi<\/jats:italic><jats:bold>ssp. nov.<\/jats:bold> A diagnostic key to all <jats:italic>Calotes<\/jats:italic> species of China is also provided.<\/jats:p>","DOI":"10.3897\/zookeys.1187.110704","type":"journal-article","created":{"date-parts":[[2023,12,20]],"date-time":"2023-12-20T15:24:00Z","timestamp":1703085840000},"page":"63-89","source":"Crossref","is-referenced-by-count":10,"title":["\ufeffTaxonomic review of the Calotes versicolor complex (Agamidae, Sauria, Squamata) in China, with description of a new species and subspecies"],"prefix":"10.3897","volume":"1187","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-3493-9468","authenticated-orcid":true,"given":"Yong","family":"Huang","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Hongyu","family":"Li","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yilin","family":"Wang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Maojin","family":"Li","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Mian","family":"Hou","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4214-9060","authenticated-orcid":true,"given":"Bo","family":"Cai","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"2258","published-online":{"date-parts":[[2023,12,20]]},"reference":[{"key":"110704_B1","doi-asserted-by":"publisher","DOI":"10.47605\/tapro.v1i1.2"},{"key":"110704_B2","doi-asserted-by":"publisher","DOI":"10.17520\/biods.2015037"},{"key":"110704_B3","doi-asserted-by":"publisher","DOI":"10.16373\/j.cnki.ahr.210020"},{"key":"110704_B4","doi-asserted-by":"publisher","DOI":"10.16373\/j.cnki.ahr.220040"},{"key":"110704_B5","doi-asserted-by":"publisher","DOI":"10.11646\/zootaxa.4317.3.10"},{"key":"110704_B6","doi-asserted-by":"publisher","DOI":"10.5962\/bhl.title.60678"},{"issue":"1","key":"110704_B7","first-page":"15","article-title":"Faunal analysis and survey on lizard in Hunan Province.","volume":"15","author":"Deng","year":"1996","journal-title":"Sichuan Journal of Zoology"},{"key":"110704_B8","doi-asserted-by":"publisher","DOI":"10.13859\/j.cjz.1997.06.014"},{"key":"110704_B9","doi-asserted-by":"publisher","DOI":"10.19711\/j.cnki.issn2310-1490.2018.03.030"},{"key":"110704_B10","doi-asserted-by":"publisher","DOI":"10.11646\/zootaxa.4638.4.1"},{"key":"110704_B11","doi-asserted-by":"publisher","DOI":"10.11646\/zootaxa.4126.2.7"},{"key":"110704_B12","doi-asserted-by":"publisher","DOI":"10.3897\/vz.71.e62787"},{"key":"110704_B13","doi-asserted-by":"publisher","DOI":"10.1177\/1934578X1100600117"},{"key":"110704_B14","doi-asserted-by":"publisher","DOI":"10.1021\/bk-1999-0734.ch008"},{"key":"110704_B15","doi-asserted-by":"publisher","DOI":"10.11646\/zootaxa.3599.3.3"},{"key":"110704_B16","doi-asserted-by":"publisher","DOI":"10.13859\/j.cjz.202201011"},{"key":"110704_B17","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0064754"},{"issue":"3","key":"110704_B18","first-page":"265","article-title":"Understanding color models: A review.","volume":"2","author":"Ibraheem","year":"2012","journal-title":"Journal of Science and Technology"},{"key":"110704_B19","author":"Kuhl","year":"1820","journal-title":"Beitrage zur Zoologie und Vergleichenden Anatomie, Erste Abtheilung."},{"key":"110704_B20","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msy096"},{"issue":"3","key":"110704_B21","first-page":"112","article-title":"Biodiversity analysis of amphibians and reptiles of Hong Kong Special Administrative Region.","volume":"19","author":"Liu","year":"2000","journal-title":"Sichuan Journal of Zoology"},{"key":"110704_B22","doi-asserted-by":"publisher","DOI":"10.3897\/herpetozoa.34.e62596"},{"key":"110704_B23","doi-asserted-by":"publisher","DOI":"10.3897\/BDJ.10.e77963"},{"key":"110704_B24","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msu300"},{"key":"110704_B25","doi-asserted-by":"publisher","DOI":"10.11646\/zootaxa.4482.3.1"},{"key":"110704_B26","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msn083"},{"key":"110704_B27","doi-asserted-by":"publisher","DOI":"10.1093\/sysbio\/sys029"},{"key":"110704_B28","author":"Shi","year":"2011"},{"key":"110704_B29","author":"Smith","year":"1935","journal-title":"The Fauna of British India including Ceylon and Burma. 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