{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,15]],"date-time":"2026-10-15T15:26:55Z","timestamp":1792078015256,"version":"4.4.0"},"reference-count":36,"publisher":"Pensoft Publishers","license":[{"start":{"date-parts":[[2026,5,22]],"date-time":"2026-05-22T00:00:00Z","timestamp":1779408000000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["ZK"],"abstract":"<jats:p>\n                    A new species of\n                    <jats:italic>Odorrana<\/jats:italic>\n                    is described from the limestone karst forest in northern Vietnam based on morphological differences and molecular divergence. Morphologically,\n                    <jats:italic>Odorrana nagao<\/jats:italic>\n                    <jats:bold>sp. nov<\/jats:bold>\n                    . is distinguishable from its congeners by the following combination of characteristics: size medium (SVL 37.4 mm in male, 47.1\u201354.7 mm in females); head longer than wide; tympanum distinct, round, ~50% of the diameter of eye; vomerine teeth present; males without external vocal sac; nuptial pad present on finger I in males; webbing formula I0\u20131II0\u20131III0\u201311\/3IV11\/3\u20131\/4V; tibio-tarsal articulation reaching the tip of snout when hindlimb adpressed along body; dorsolateral conical spines present; dorsolateral fold present; dorsum moss-green with brown mottling; ventral surface cream with dark brown mottling. Phylogenetic analyses of mitochondrial 16S rRNA and COI gene sequences supported the new species as a sister taxon to an unresolved clade consisting of\n                    <jats:italic>Odorrana calciphila<\/jats:italic>\n                    ,\n                    <jats:italic>O. concelata<\/jats:italic>\n                    ,\n                    <jats:italic>O. feii<\/jats:italic>\n                    ,\n                    <jats:italic>O. liboensis<\/jats:italic>\n                    and\n                    <jats:italic>O. lipuensis<\/jats:italic>\n                    (UFB = 98, PP = 1). The uncorrected p-distances of 16S rRNA and COI genes between the new species and its closest congeners were 2.33% and 6.33%, respectively.\n                  <\/jats:p>","DOI":"10.3897\/zookeys.1280.192981","type":"journal-article","created":{"date-parts":[[2026,5,22]],"date-time":"2026-05-22T13:43:48Z","timestamp":1779457428000},"page":"245-263","source":"Crossref","is-referenced-by-count":2,"title":["A new species of Odorrana (Anura, Ranidae) from the limestone karst forest of northern Vietnam"],"prefix":"10.3897","volume":"1280","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-5158-4526","authenticated-orcid":true,"given":"Cuong The","family":"Pham","sequence":"first","affiliation":[{"name":"Institute of Biology, Vietnam Academy of Science and Technology, Hanoi, Vietnam"},{"name":"Graduate University of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-0709-974X","authenticated-orcid":true,"given":"Chung Van","family":"Hoang","sequence":"additional","affiliation":[{"name":"Institute of Biology, Vietnam Academy of Science and Technology, Hanoi, Vietnam"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0001-8322-010X","authenticated-orcid":true,"given":"Mai Hong Thi","family":"Nguyen","sequence":"additional","affiliation":[{"name":"Institute of Biology, Vietnam Academy of Science and Technology, Hanoi, Vietnam"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-6601-0880","authenticated-orcid":true,"given":"Truong Quang","family":"Nguyen","sequence":"additional","affiliation":[{"name":"Institute of Biology, Vietnam Academy of Science and Technology, Hanoi, Vietnam"},{"name":"Graduate University of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-5471-965X","authenticated-orcid":true,"given":"Hai Ngoc","family":"Ngo","sequence":"additional","affiliation":[{"name":"Institute of Theoretical and Applied Research, Duy Tan University, Hanoi, Vietnam"},{"name":"School of Medicine and Pharmacy, Duy Tan University, Da Nang, Vietnam"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5925-7797","authenticated-orcid":true,"given":"Dzung Trung","family":"Le","sequence":"additional","affiliation":[{"name":"Faculty of Biology, Thai Nguyen University of Education, Thai Nguyen, Vietnam"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4797-609X","authenticated-orcid":true,"given":"Thomas","family":"Ziegler","sequence":"additional","affiliation":[{"name":"Cologne Zoo, Cologne, Germany"},{"name":"Institute of Zoology, University of Cologne, Cologne, Germany"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"2258","published-online":{"date-parts":[[2026,5,22]]},"reference":[{"key":"192981_B1","doi-asserted-by":"publisher","DOI":"10.11646\/zootaxa.5124.4.4"},{"key":"192981_B2","doi-asserted-by":"publisher","DOI":"10.1111\/j.1755-0998.2011.03090.x"},{"key":"192981_B3","doi-asserted-by":"publisher","DOI":"10.1641\/0006-3568(2006)56[733:LKOSAI]2.0.CO;2"},{"key":"192981_B4","doi-asserted-by":"publisher","DOI":"10.1038\/nmeth.2109"},{"key":"192981_B5","unstructured":"Fei L, Hu SQ, Ye CY, Huang YZ (2009) Fauna Sinica. 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