{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,2]],"date-time":"2026-10-02T10:59:57Z","timestamp":1790938797297,"version":"4.1.0"},"reference-count":49,"publisher":"Pensoft Publishers","license":[{"start":{"date-parts":[[2026,10,2]],"date-time":"2026-10-02T00:00:00Z","timestamp":1790899200000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["ZK"],"abstract":"<jats:p>\n                    An integrative taxonomic study combining morphological and molecular data revealed two new species of giant pill-millipedes of the genus\n                    <jats:italic>Zephronia<\/jats:italic>\n                    Gray, 1832:\n                    <jats:italic>Zephronia tamarindina<\/jats:italic>\n                    Wilaisorn &amp; Srisonchai,\n                    <jats:bold>sp. nov<\/jats:bold>\n                    . and\n                    <jats:italic>Zephronia triforma<\/jats:italic>\n                    Wilaisorn &amp; Srisonchai,\n                    <jats:bold>sp. nov<\/jats:bold>\n                    ., both endemic to provinces in the upper central region of Thailand. In addition,\n                    <jats:italic>Zephronia dawydoffi<\/jats:italic>\n                    Attems, 1953 and\n                    <jats:italic>Zephronia laotica<\/jats:italic>\n                    Wesener, 2019 are recorded from Thailand for the first time. The new species exhibit distinctive colouration patterns, and scanning electron microscopy revealed unique morphological characters for each taxon. Analysis of the 658 bp DNA barcoding region of the mitochondrial\n                    <jats:italic>COI<\/jats:italic>\n                    gene showed interspecific genetic distances ranging from 8.68\u201321.92% for\n                    <jats:italic>Z. dawydoffi<\/jats:italic>\n                    , 12.79\u201322.98% for\n                    <jats:italic>Z. laotica<\/jats:italic>\n                    , 10.81\u201322.07% for\n                    <jats:italic>Z. tamarindina<\/jats:italic>\n                    <jats:bold>sp. nov<\/jats:bold>\n                    ., and 10.81\u201322.83% for\n                    <jats:italic>Z. triforma<\/jats:italic>\n                    <jats:bold>sp. nov<\/jats:bold>\n                    ., in comparison with closely related species from Thailand, Vietnam, Laos, and Cambodia. Phylogenetic analyses placed all four species within the\n                    <jats:italic>Zephronia<\/jats:italic>\n                    s.s. clade, characterised by the T\u00f6m\u00f6sv\u00e1ry\u2019s organ located outside the antennal groove. However, relationships among other clades remained unresolved due to interspersion with species of\n                    <jats:italic>Cryxus<\/jats:italic>\n                    Leach, 1814,\n                    <jats:italic>Prionobelum<\/jats:italic>\n                    Verhoeff, 1924, and\n                    <jats:italic>Sphaerobelum<\/jats:italic>\n                    Verhoeff, 1924.\n                    <jats:italic>Zephronia<\/jats:italic>\n                    was recovered as non-monophyletic, indicating that the current generic limits within Zephroniinae require reassessment using multi-locus data. These results expand our understanding of giant pill-millipede diversity and highlight the upper central Thai region and Southeast Asia more broadly as a significant biodiversity hotspot.\n                  <\/jats:p>","DOI":"10.3897\/zookeys.1294.205518","type":"journal-article","created":{"date-parts":[[2026,10,2]],"date-time":"2026-10-02T10:40:32Z","timestamp":1790937632000},"page":"1-59","source":"Crossref","is-referenced-by-count":0,"title":["Two new species and two new country records of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand, based on integrative taxonomy (Diplopoda, Sphaerotheriida, Zephroniidae)"],"prefix":"10.3897","volume":"1294","author":[{"given":"Chalintorn","family":"Wilaisorn","sequence":"first","affiliation":[{"name":"Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, Thailand"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ruttapon","family":"Srisonchai","sequence":"additional","affiliation":[{"name":"Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, Thailand"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wibhu","family":"Kutanan","sequence":"additional","affiliation":[{"name":"Department of Biology, Faculty of Science, Naresuan University, Phitsanulok, Thailand"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Nisarat","family":"Tungpairojwong","sequence":"additional","affiliation":[{"name":"Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, Thailand"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"2258","published-online":{"date-parts":[[2026,10,2]]},"reference":[{"key":"205518_B1","unstructured":"American Veterinary Medical Association (2020) AVMA Guidelines for the Euthanasia of Animals: 2020 Edition. https:\/\/www.avma.org\/sites\/default\/files\/2020-02\/Guidelines-on-Euthanasia-2020.pdf [accessed 16 January 2025]"},{"key":"205518_B2","unstructured":"Attems C (1953) Myriopoden von Indochina, Expedition von Dr. C. 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