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Using a whole-brain searchlight, we observed cross-feature decoding not in retinotopic cortex, but bilaterally in anterior inferior temporal cortex (aIT), temporal pole (TP), and frontal cortex (FC). Only TP and aIT had response patterns to concept shapes matching color-space geometry, and only PF had responses that decode behavioral errors. Two additional imaging experiments showed that color-associated versus non-color-associated shapes, and color-shape combinations congruent versus incongruent with the concepts, were distinguished only by extrastriate visual cortex, suggesting extrastriate cortex underpins color perception and perceptual feature binding. 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