{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"institution":[{"name":"bioRxiv"}],"indexed":{"date-parts":[[2026,5,21]],"date-time":"2026-05-21T16:05:41Z","timestamp":1779379541873,"version":"3.53.1"},"posted":{"date-parts":[[2026,5,18]]},"group-title":"Neuroscience","reference-count":106,"publisher":"openRxiv","license":[{"start":{"date-parts":[[2026,5,18]],"date-time":"2026-05-18T00:00:00Z","timestamp":1779062400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by-nc\/4.0\/"}],"funder":[{"name":"NIH R01","award":["R01NS115862 to Aaron L. Wong"],"award-info":[{"award-number":["R01NS115862 to Aaron L. Wong"]}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"accepted":{"date-parts":[[2026,5,18]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                <jats:p>A hallmark of human intelligence is the ability to use tools. Yet the cognitive processes supporting this ability remain debated. One contemporary view holds that mechanical reasoning is central for tool use, especially in the case of tools with which we have no prior experience. However, previous support for the role of mechanical reasoning often relies on circular logic, wherein poor performance on novel tool-use tasks is taken as evidence that impaired mechanical reasoning causes tool-use deficits in limb apraxia. To address this limitation, we independently assessed mechanical reasoning and novel tool use in separate tasks in individuals with limb apraxia, and compared their performance to individuals without apraxia. We also examined whether these two abilities are similarly associated with other cognitive abilities including motor imagery, mental rotation of non-body objects, general reasoning, and spatial working memory. Finally, we explored brain-behavior relationships using support vector regression lesion-symptom mapping. Our behavioral and imaging data together showed that mechanical reasoning does not underlie novel tool-use deficits in apraxia. Graphical analysis further revealed that novel tool use and mechanical reasoning loaded onto distinct latent clusters: novel tool use was strongly associated with other praxis abilities yet separable from cognitive abilities that require reasoning and mental simulation, whereas mechanical reasoning was primarily linked to other high-level reasoning abilities but not tool use. These findings challenge the notion that mechanical reasoning is central to tool-use ability, and instead suggest that tool use is more likely to be an intuitive or automatic process.<\/jats:p>","DOI":"10.64898\/2026.05.14.724638","type":"posted-content","created":{"date-parts":[[2026,5,18]],"date-time":"2026-05-18T19:10:14Z","timestamp":1779131414000},"source":"Crossref","is-referenced-by-count":0,"title":["Novel tool use does not depend on mechanical reasoning: evidence from apraxia"],"prefix":"10.64898","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-3865-0805","authenticated-orcid":false,"given":"Yue","family":"Du","sequence":"first","affiliation":[{"name":"Department of Neurology, Johns Hopkins University, Baltimore, MD, USA"},{"name":"Department of Kinesiology and Sport Management, Texas A&M University, College Station, TX, USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Simon","family":"Thibault","sequence":"additional","affiliation":[{"name":"Jefferson Moss Rehabilitation Research Institute, Thomas Jefferson University, Philadelphia, PA, USA"},{"name":"Centre de Recherche en Psychologie et Neurosciences, Aix-Marseille Universit\u00e9"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"John","family":"Yates","sequence":"additional","affiliation":[{"name":"Jefferson Moss Rehabilitation Research Institute, Thomas Jefferson University, Philadelphia, PA, USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Laurel J.","family":"Buxbaum","sequence":"additional","affiliation":[{"name":"Jefferson Moss Rehabilitation Research Institute, Thomas Jefferson University, Philadelphia, PA, USA"},{"name":"Department of Rehabilitation Medicine, Sidney Kimmel Medical College, Thomas Jefferson University, Philadelphia, PA, USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"John W.","family":"Krakauer","sequence":"additional","affiliation":[{"name":"Department of Neurology, Johns Hopkins University, Baltimore, MD, USA"},{"name":"Department of Neuroscience, Johns Hopkins University, Baltimore, MD, USA"},{"name":"Department of Physical Medicine and Rehabilitation, Johns Hopkins University, Baltimore, MD, USA"},{"name":"The Santa Fe Institute, Santa Fe, NM, USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Aaron L.","family":"Wong","sequence":"additional","affiliation":[{"name":"Jefferson Moss Rehabilitation Research Institute, Thomas Jefferson University, Philadelphia, PA, USA"},{"name":"Department of Rehabilitation Medicine, Sidney Kimmel Medical College, Thomas Jefferson University, Philadelphia, PA, USA"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"54368","reference":[{"key":"2026052108153439000_2026.05.14.724638v1.1","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1912341117"},{"key":"2026052108153439000_2026.05.14.724638v1.2","doi-asserted-by":"publisher","DOI":"10.1037\/0096-3445.135.3.327"},{"issue":"3","key":"2026052108153439000_2026.05.14.724638v1.3","doi-asserted-by":"crossref","first-page":"143","DOI":"10.1155\/2007\/241670","article-title":"Relationship between object-related gestures and the fractionated object knowledge system","volume":"18","year":"2007","journal-title":"Behavioural Neurology"},{"key":"2026052108153439000_2026.05.14.724638v1.4","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1306572110"},{"key":"2026052108153439000_2026.05.14.724638v1.5","doi-asserted-by":"publisher","DOI":"10.1080\/02643294.2022.2071152"},{"key":"2026052108153439000_2026.05.14.724638v1.6","doi-asserted-by":"crossref","first-page":"119","DOI":"10.1016\/j.cortex.2016.06.007","article-title":"Tool use disorders in neurodegenerative diseases: Roles of semantic memory and technical reasoning","volume":"82","year":"2016","journal-title":"Cortex"},{"issue":"3","key":"2026052108153439000_2026.05.14.724638v1.7","doi-asserted-by":"crossref","first-page":"409","DOI":"10.1111\/jnp.12121","article-title":"Tool use in neurodegenerative diseases: Planning or technical reasoning?","volume":"12","year":"2018","journal-title":"Journal of Neuropsychology"},{"key":"2026052108153439000_2026.05.14.724638v1.8","first-page":"473","article-title":"Tool use disorders after left brain damage","volume":"5","year":"2014","journal-title":"Frontiers in Psychology"},{"key":"2026052108153439000_2026.05.14.724638v1.9","doi-asserted-by":"publisher","DOI":"10.3758\/BF03196585"},{"key":"2026052108153439000_2026.05.14.724638v1.10","first-page":"191","article-title":"An investigation of limb apraxia regarding the validity of current assessment procedures","volume":"22","year":"1994","journal-title":"Clinical Aphasiology"},{"key":"2026052108153439000_2026.05.14.724638v1.11","doi-asserted-by":"publisher","DOI":"10.1177\/1073191112446655"},{"key":"2026052108153439000_2026.05.14.724638v1.12","doi-asserted-by":"publisher","DOI":"10.1016\/j.cognition.2006.12.010"},{"key":"2026052108153439000_2026.05.14.724638v1.13","doi-asserted-by":"crossref","unstructured":"Buxbaum, L. J. (2017). Learning, remembering, and predicting how to use tools: Distributed neurocognitive mechanisms: Comment on Osiurak and Badets (2016).","DOI":"10.1037\/rev0000051"},{"key":"2026052108153439000_2026.05.14.724638v1.14","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuropsychologia.2004.09.006"},{"key":"2026052108153439000_2026.05.14.724638v1.15","doi-asserted-by":"crossref","first-page":"349","DOI":"10.1016\/B978-0-444-63622-5.00017-6","article-title":"Limb apraxia and the left parietal lobe","volume":"151","year":"2018","journal-title":"Handbook of Clinical Neurology"},{"key":"2026052108153439000_2026.05.14.724638v1.16","doi-asserted-by":"publisher","DOI":"10.1016\/S0093-934X(02)00014-7"},{"key":"2026052108153439000_2026.05.14.724638v1.17","doi-asserted-by":"publisher","DOI":"10.1093\/brain\/awu111"},{"key":"2026052108153439000_2026.05.14.724638v1.18","doi-asserted-by":"publisher","DOI":"10.1093\/brain\/117.5.1093"},{"key":"2026052108153439000_2026.05.14.724638v1.19","doi-asserted-by":"publisher","DOI":"10.1016\/0010-0285(75)90003-1"},{"key":"2026052108153439000_2026.05.14.724638v1.20","unstructured":"Corporation, P. (1990). The Mechanical Reasoning Test of the Differential Aptitudes Test. Appendices New York."},{"key":"2026052108153439000_2026.05.14.724638v1.21","unstructured":"Corsi, P. M. , & Michael, P. (1972). Human memory and the medial temporal region of the brain."},{"key":"2026052108153439000_2026.05.14.724638v1.22","unstructured":"Cronbach, L. J. (1970). Essentials of psychological testing."},{"key":"2026052108153439000_2026.05.14.724638v1.23","doi-asserted-by":"publisher","DOI":"10.1016\/0004-3702(84)90037-7"},{"issue":"5","key":"2026052108153439000_2026.05.14.724638v1.24","doi-asserted-by":"crossref","first-page":"446","DOI":"10.1038\/s41562-019-0567-9","article-title":"Causal understanding is not necessary for the improvement of culturally evolving technology","volume":"3","year":"2019","journal-title":"Nature Human Behaviour"},{"key":"2026052108153439000_2026.05.14.724638v1.25","doi-asserted-by":"crossref","first-page":"153","DOI":"10.1016\/j.neuropsychologia.2016.04.020","article-title":"Perceptual decisions regarding object manipulation are selectively impaired in apraxia or when tDCS is applied over the left IPL","volume":"86","year":"2016","journal-title":"Neuropsychologia"},{"key":"2026052108153439000_2026.05.14.724638v1.26","unstructured":"Evans, J. S. B. , & Over, D. E. (2013). Rationality and reasoning. Psychology Press. https:\/\/api.taylorfrancis.com\/content\/books\/mono\/download?identifierName=doi&identifierValue=10.4324\/9780203027677&type=googlepdf"},{"issue":"3","key":"2026052108153439000_2026.05.14.724638v1.27","doi-asserted-by":"crossref","first-page":"249","DOI":"10.1037\/neu0000402","article-title":"Tool use in left-brain-damaged patients: Difficulties in reasoning but not in estimating the physical properties of objects","volume":"32","year":"2018","journal-title":"Neuropsychology"},{"key":"2026052108153439000_2026.05.14.724638v1.28","doi-asserted-by":"publisher","DOI":"10.1080\/02643294.2022.2106126"},{"key":"2026052108153439000_2026.05.14.724638v1.29","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1610344113"},{"key":"2026052108153439000_2026.05.14.724638v1.30","unstructured":"Fitts, P. M. , & Posner, M. I . (1967). Human performance. Brooks and Cole."},{"key":"2026052108153439000_2026.05.14.724638v1.31","doi-asserted-by":"publisher","DOI":"10.1093\/biostatistics\/kxm045"},{"key":"2026052108153439000_2026.05.14.724638v1.32","doi-asserted-by":"publisher","DOI":"10.1037\/0096-1523.15.2.372"},{"issue":"2","key":"2026052108153439000_2026.05.14.724638v1.33","doi-asserted-by":"crossref","first-page":"176","DOI":"10.1136\/jnnp.61.2.176","article-title":"Defective imitation of gestures in patients with damage in the left or right hemispheres. Journal of Neurology","volume":"61","year":"1996","journal-title":"Neurosurgery & Psychiatry"},{"issue":"5","key":"2026052108153439000_2026.05.14.724638v1.34","first-page":"453","article-title":"Apraxia","volume":"4","year":"2013","journal-title":"Wiley Interdisciplinary Reviews: Cognitive Science"},{"key":"2026052108153439000_2026.05.14.724638v1.35","doi-asserted-by":"crossref","unstructured":"Goldenberg, G. (2013b). Apraxia: The cognitive side of motor control. Oup Oxford.","DOI":"10.1093\/acprof:oso\/9780199591510.001.0001"},{"key":"2026052108153439000_2026.05.14.724638v1.36","doi-asserted-by":"publisher","DOI":"10.1016\/S0028-3932(97)00165-6"},{"key":"2026052108153439000_2026.05.14.724638v1.37","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuropsychologia.2015.05.017"},{"key":"2026052108153439000_2026.05.14.724638v1.38","doi-asserted-by":"publisher","DOI":"10.1093\/brain\/awp080"},{"key":"2026052108153439000_2026.05.14.724638v1.39","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0174035"},{"key":"2026052108153439000_2026.05.14.724638v1.40","doi-asserted-by":"publisher","DOI":"10.1016\/j.tics.2021.01.006"},{"key":"2026052108153439000_2026.05.14.724638v1.41","doi-asserted-by":"publisher","DOI":"10.1016\/j.cognition.2016.08.012"},{"key":"2026052108153439000_2026.05.14.724638v1.42","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuropsychologia.2004.07.015"},{"issue":"5","key":"2026052108153439000_2026.05.14.724638v1.43","doi-asserted-by":"crossref","first-page":"1084","DOI":"10.1037\/0278-7393.18.5.1084","article-title":"Mental animation: Inferring motion from static displays of mechanical systems. Journal of Experimental Psychology: Learning","volume":"18","year":"1992","journal-title":"Memory, and Cognition"},{"key":"2026052108153439000_2026.05.14.724638v1.44","doi-asserted-by":"publisher","DOI":"10.1016\/j.tics.2004.04.001"},{"key":"2026052108153439000_2026.05.14.724638v1.45","doi-asserted-by":"publisher","DOI":"10.1016\/0010-0285(88)90019-9"},{"issue":"4","key":"2026052108153439000_2026.05.14.724638v1.46","doi-asserted-by":"crossref","first-page":"333","DOI":"10.1207\/s15427633scc0504_3","article-title":"The role of gestures in mental animation","volume":"5","year":"2005","journal-title":"Spatial Cognition and Computation"},{"key":"2026052108153439000_2026.05.14.724638v1.47","doi-asserted-by":"publisher","DOI":"10.1016\/j.cortex.2011.10.010"},{"key":"2026052108153439000_2026.05.14.724638v1.48","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.96.16.9444"},{"key":"2026052108153439000_2026.05.14.724638v1.49","doi-asserted-by":"crossref","unstructured":"Jarry, C. , Osiurak, F. , Baumard, J. , Lesourd, M. , Etcharry-Bouyx, F. , Chauvir\u00e9, V. , & Le Gall, D. (2015). Mechanical problem-solving and imitation of meaningless postures in left brain damaged patients: Two sides of the same coin ?","DOI":"10.1016\/j.cortex.2014.08.021"},{"key":"2026052108153439000_2026.05.14.724638v1.50","doi-asserted-by":"publisher","DOI":"10.1016\/j.cortex.2013.02.011"},{"key":"2026052108153439000_2026.05.14.724638v1.51","doi-asserted-by":"publisher","DOI":"10.1016\/j.tics.2003.12.002"},{"key":"2026052108153439000_2026.05.14.724638v1.52","unstructured":"Kahneman, D. (2011). Thinking, fast and slow. macmillan. https:\/\/books.google.com\/books?hl=en&lr=&id=SHvzzuCnuv8C&oi=fnd&pg=PP2&dq=Daniel+Kahneman.&ots=NUpkMC6mFD&sig=v6IFhYLdiMFCsVD8zJw6Uu9Yrsc"},{"key":"2026052108153439000_2026.05.14.724638v1.53","unstructured":"Kertesz, A. (2007). Western aphasia battery\u2013revised. https:\/\/psycnet.apa.org\/doiLanding?doi=10.1037%2Ft15168-000"},{"key":"2026052108153439000_2026.05.14.724638v1.54","doi-asserted-by":"publisher","DOI":"10.1017\/S0048577298001516"},{"issue":"4","key":"2026052108153439000_2026.05.14.724638v1.55","doi-asserted-by":"crossref","first-page":"952","DOI":"10.1037\/0278-7393.19.4.952","article-title":"Intuitive physics in action and judgment: The development of knowledge about projectile motion. Journal of Experimental Psychology: Learning","volume":"19","year":"1993","journal-title":"Memory, and Cognition"},{"key":"2026052108153439000_2026.05.14.724638v1.56","doi-asserted-by":"publisher","DOI":"10.1016\/j.tics.2017.06.002"},{"issue":"2","key":"2026052108153439000_2026.05.14.724638v1.57","doi-asserted-by":"crossref","first-page":"198","DOI":"10.1111\/jnp.12140","article-title":"Mechanical knowledge does matter to tool use even when assessed with a non-production task: Evidence from left brain-damaged patients","volume":"13","year":"2019","journal-title":"Journal of Neuropsychology"},{"issue":"1","key":"2026052108153439000_2026.05.14.724638v1.58","doi-asserted-by":"crossref","first-page":"552","DOI":"10.1038\/s42003-025-07923-1","article-title":"Neurocognitive mechanisms underlying action tool knowledge tasks: Specificity of tool-tool compared to hand-tool compatibility tasks","volume":"8","year":"2025","journal-title":"Communications Biology"},{"key":"2026052108153439000_2026.05.14.724638v1.59","doi-asserted-by":"crossref","unstructured":"Liu, S. , Karakose-Akbiyik, S. , Caramazza, A. , Sotero, F. D. , Buxbaum, L. J. , & Wong, A. L . (2025). Dissociation between physical reasoning and tool use in individuals with left hemisphere brain damage. bioRxiv, 2025\u201311.","DOI":"10.1101\/2025.11.20.688863"},{"key":"2026052108153439000_2026.05.14.724638v1.60","doi-asserted-by":"crossref","unstructured":"Mangalam, M. , Fragaszy, D. M. , Wagman, J. B. , Day, B. M. , Kelty-Stephen, D. G. , Bongers, R. M. , Stout, D. W. , & Osiurak, F . (2022). On the psychological origins of tool use. Neuroscience & Biobehavioral Reviews, 104521.","DOI":"10.1016\/j.neubiorev.2022.104521"},{"key":"2026052108153439000_2026.05.14.724638v1.61","doi-asserted-by":"publisher","DOI":"10.1080\/09541440701394624"},{"key":"2026052108153439000_2026.05.14.724638v1.62","doi-asserted-by":"publisher","DOI":"10.1186\/s41235-020-00224-7"},{"key":"2026052108153439000_2026.05.14.724638v1.63","first-page":"1","article-title":"A dedicated mental resource for intuitive physics","volume":"27","year":"2024","journal-title":"Iscience"},{"key":"2026052108153439000_2026.05.14.724638v1.64","doi-asserted-by":"publisher","DOI":"10.1016\/j.jneumeth.2013.10.024"},{"key":"2026052108153439000_2026.05.14.724638v1.65","doi-asserted-by":"publisher","DOI":"10.1016\/j.bandl.2009.12.003"},{"key":"2026052108153439000_2026.05.14.724638v1.66","doi-asserted-by":"publisher","DOI":"10.1212\/WNL.49.2.474"},{"key":"2026052108153439000_2026.05.14.724638v1.67","first-page":"310","article-title":"The neural basis of human tool use","volume":"5","year":"2014","journal-title":"Frontiers in Psychology"},{"key":"2026052108153439000_2026.05.14.724638v1.68","doi-asserted-by":"publisher","DOI":"10.1037\/rev0000027"},{"key":"2026052108153439000_2026.05.14.724638v1.69","doi-asserted-by":"publisher","DOI":"10.1016\/j.cortex.2008.06.013"},{"key":"2026052108153439000_2026.05.14.724638v1.70","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuropsychologia.2013.06.017"},{"issue":"12","key":"2026052108153439000_2026.05.14.724638v1.71","doi-asserted-by":"crossref","first-page":"1643","DOI":"10.1038\/s41562-021-01159-9","article-title":"Technical reasoning is important for cumulative technological culture","volume":"5","year":"2021","journal-title":"Nature Human Behaviour"},{"key":"2026052108153439000_2026.05.14.724638v1.72","doi-asserted-by":"crossref","unstructured":"Osiurak, F. , & Reynaud, E . (2020). The elephant in the room: What matters cognitively in cumulative technological culture. Behavioral and Brain Sciences, 43.","DOI":"10.1017\/S0140525X20000291"},{"issue":"4","key":"2026052108153439000_2026.05.14.724638v1.73","first-page":"fcab263","article-title":"Pantomime of tool use: Looking beyond apraxia","volume":"3","year":"2021","journal-title":"Brain Communications"},{"key":"2026052108153439000_2026.05.14.724638v1.74","doi-asserted-by":"publisher","DOI":"10.1037\/0096-1523.20.4.709"},{"key":"2026052108153439000_2026.05.14.724638v1.75","doi-asserted-by":"publisher","DOI":"10.3758\/s13428-018-01193-y"},{"key":"2026052108153439000_2026.05.14.724638v1.76","doi-asserted-by":"publisher","DOI":"10.1093\/brain\/118.1.227"},{"key":"2026052108153439000_2026.05.14.724638v1.77","unstructured":"R Core Team . (2013). R: A language and environment for statistical computing [Computer software]. R Foundation for Statistical Computing. http:\/\/www.R-project.org\/."},{"key":"2026052108153439000_2026.05.14.724638v1.78","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuropsychologia.2011.04.017"},{"key":"2026052108153439000_2026.05.14.724638v1.79","doi-asserted-by":"publisher","DOI":"10.1080\/13506280344000428"},{"key":"2026052108153439000_2026.05.14.724638v1.80","doi-asserted-by":"publisher","DOI":"10.1016\/j.neubiorev.2016.03.009"},{"key":"2026052108153439000_2026.05.14.724638v1.81","doi-asserted-by":"crossref","first-page":"10","DOI":"10.1007\/s00426-013-0481-0","article-title":"Tool characteristics in imagery of tool actions","volume":"78","year":"2014","journal-title":"Psychological Research"},{"key":"2026052108153439000_2026.05.14.724638v1.82","unstructured":"Rothi, L. J. G. , & Heilman, K. M. (2014). Apraxia: The neuropsychology of action. Psychology Press. https:\/\/api.taylorfrancis.com\/content\/books\/mono\/download?identifierName=doi&identifierValue=10.4324\/9781315804545&type=googlepdf"},{"issue":"4","key":"2026052108153439000_2026.05.14.724638v1.83","doi-asserted-by":"crossref","first-page":"293","DOI":"10.1123\/apaq.10.4.293","article-title":"Memory processes and gestural performance in apraxia","volume":"10","year":"1993","journal-title":"Adapted Physical Activity Quarterly"},{"key":"2026052108153439000_2026.05.14.724638v1.84","doi-asserted-by":"publisher","DOI":"10.1007\/978-1-4471-0109-3_13"},{"key":"2026052108153439000_2026.05.14.724638v1.85","doi-asserted-by":"publisher","DOI":"10.1016\/S0010-9452(08)70626-9"},{"key":"2026052108153439000_2026.05.14.724638v1.86","doi-asserted-by":"publisher","DOI":"10.1006\/cogp.1996.0006"},{"issue":"4","key":"2026052108153439000_2026.05.14.724638v1.87","doi-asserted-by":"crossref","first-page":"457","DOI":"10.1207\/s15516709cog2004_1","article-title":"Shuttling between depictive models and abstract rules: Induction and fallback","volume":"20","year":"1996","journal-title":"Cognitive Science"},{"issue":"1","key":"2026052108153439000_2026.05.14.724638v1.88","doi-asserted-by":"crossref","first-page":"116","DOI":"10.1037\/0278-7393.25.1.116","article-title":"Inferences through imagined actions: Knowing by simulated doing. Journal of Experimental Psychology: Learning","volume":"25","year":"1999","journal-title":"Memory, and Cognition"},{"issue":"6","key":"2026052108153439000_2026.05.14.724638v1.89","doi-asserted-by":"crossref","first-page":"1655","DOI":"10.1037\/0278-7393.26.6.1655","article-title":"Tool use and the effect of action on the imagination. Journal of Experimental Psychology: Learning","volume":"26","year":"2000","journal-title":"Memory, and Cognition"},{"key":"2026052108153439000_2026.05.14.724638v1.90","doi-asserted-by":"publisher","DOI":"10.1080\/02643290342000348"},{"key":"2026052108153439000_2026.05.14.724638v1.91","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuroimage.2025.121073"},{"key":"2026052108153439000_2026.05.14.724638v1.92","doi-asserted-by":"publisher","DOI":"10.1126\/science.273.5281.1564"},{"key":"2026052108153439000_2026.05.14.724638v1.93","doi-asserted-by":"publisher","DOI":"10.1007\/s004150200070"},{"key":"2026052108153439000_2026.05.14.724638v1.94","doi-asserted-by":"publisher","DOI":"10.1016\/j.cortex.2021.10.002"},{"key":"2026052108153439000_2026.05.14.724638v1.95","doi-asserted-by":"publisher","DOI":"10.1523\/JNEUROSCI.0692-25.2025"},{"key":"2026052108153439000_2026.05.14.724638v1.96","doi-asserted-by":"publisher","DOI":"10.1016\/S0278-2626(02)00570-5"},{"key":"2026052108153439000_2026.05.14.724638v1.97","doi-asserted-by":"publisher","DOI":"10.1037\/0096-1523.24.3.830"},{"key":"2026052108153439000_2026.05.14.724638v1.98","doi-asserted-by":"publisher","DOI":"10.1016\/j.tics.2017.05.012"},{"key":"2026052108153439000_2026.05.14.724638v1.99","doi-asserted-by":"publisher","DOI":"10.1017\/S0140525X11001452"},{"issue":"2","key":"2026052108153439000_2026.05.14.724638v1.100","doi-asserted-by":"crossref","first-page":"220","DOI":"10.1016\/j.plrev.2013.11.005","article-title":"Action semantics: A unifying conceptual framework for the selective use of multimodal and modality-specific object knowledge","volume":"11","year":"2014","journal-title":"Physics of Life Reviews"},{"key":"2026052108153439000_2026.05.14.724638v1.101","first-page":"299","article-title":"Effect of biomechanical constraints in the hand laterality judgment task: Where does it come from?","volume":"6","year":"2012","journal-title":"Frontiers in Human Neuroscience"},{"key":"2026052108153439000_2026.05.14.724638v1.102","doi-asserted-by":"publisher","DOI":"10.1016\/j.cortex.2015.01.007"},{"issue":"1","key":"2026052108153439000_2026.05.14.724638v1.103","first-page":"3","article-title":"Assessing mechanistic reasoning: Supporting systems tracing","volume":"9","year":"2019","journal-title":"Journal of Pre-College Engineering Education Research (J-PEER"},{"key":"2026052108153439000_2026.05.14.724638v1.104","doi-asserted-by":"publisher","DOI":"10.1016\/S0010-0277(98)00032-8"},{"key":"2026052108153439000_2026.05.14.724638v1.105","doi-asserted-by":"publisher","DOI":"10.3758\/BF03194431"},{"key":"2026052108153439000_2026.05.14.724638v1.106","doi-asserted-by":"publisher","DOI":"10.1007\/978-3-540-25931-2_29"}],"container-title":[],"original-title":[],"link":[{"URL":"https:\/\/syndication.highwire.org\/content\/doi\/10.64898\/2026.05.14.724638","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,5,21]],"date-time":"2026-05-21T15:15:54Z","timestamp":1779376554000},"score":1,"resource":{"primary":{"URL":"http:\/\/biorxiv.org\/lookup\/doi\/10.64898\/2026.05.14.724638"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,5,18]]},"references-count":106,"URL":"https:\/\/doi.org\/10.64898\/2026.05.14.724638","relation":{},"subject":[],"published":{"date-parts":[[2026,5,18]]},"subtype":"preprint"}}