{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"institution":[{"name":"bioRxiv"}],"indexed":{"date-parts":[[2026,3,8]],"date-time":"2026-03-08T03:56:03Z","timestamp":1772942163758,"version":"3.50.1"},"posted":{"date-parts":[[2026,3,4]]},"group-title":"Evolutionary Biology","reference-count":102,"publisher":"openRxiv","license":[{"start":{"date-parts":[[2026,3,4]],"date-time":"2026-03-04T00:00:00Z","timestamp":1772582400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0\/"}],"funder":[{"award":["757451"],"award-info":[{"award-number":["757451"]}],"id":[{"id":"https:\/\/ror.org\/0472cxd90","id-type":"ROR","asserted-by":"publisher"}]},{"award":["101132305"],"award-info":[{"award-number":["101132305"]}],"id":[{"id":"https:\/\/ror.org\/0472cxd90","id-type":"ROR","asserted-by":"publisher"}]},{"award":["2019-04452"],"award-info":[{"award-number":["2019-04452"]}],"id":[{"id":"https:\/\/ror.org\/03zttf063","id-type":"ROR","asserted-by":"publisher"}]},{"award":["2023-04532"],"award-info":[{"award-number":["2023-04532"]}],"id":[{"id":"https:\/\/ror.org\/03zttf063","id-type":"ROR","asserted-by":"publisher"}]},{"award":["2022-06725"],"award-info":[{"award-number":["2022-06725"]}],"id":[{"id":"https:\/\/ror.org\/03zttf063","id-type":"ROR","asserted-by":"publisher"}]},{"id":[{"id":"https:\/\/ror.org\/057dvs720","id-type":"ROR","asserted-by":"publisher"}]},{"name":"Nilsson-Ehle foundation"},{"DOI":"10.13039\/501100002805","name":"Carl Tryggers stiftelse f\u00f6r vetenskaplig forskning","doi-asserted-by":"crossref","award":["CTS21:1471"],"award-info":[{"award-number":["CTS21:1471"]}],"id":[{"id":"10.13039\/501100002805","id-type":"DOI","asserted-by":"crossref"}]},{"id":[{"id":"https:\/\/ror.org\/01bs3p986","id-type":"ROR","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100004063","name":"Knut and Alice Wallenberg Foundation","doi-asserted-by":"crossref","award":["KAW 2020.0239"],"award-info":[{"award-number":["KAW 2020.0239"]}],"id":[{"id":"10.13039\/501100004063","id-type":"DOI","asserted-by":"crossref"}]},{"DOI":"10.13039\/501100004063","name":"Knut and Alice Wallenberg Foundation","doi-asserted-by":"crossref","award":["KAW 2017.0003"],"award-info":[{"award-number":["KAW 2017.0003"]}],"id":[{"id":"10.13039\/501100004063","id-type":"DOI","asserted-by":"crossref"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"accepted":{"date-parts":[[2026,3,4]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                <jats:p>\n                  The breakdown of distyly, a polymorphism that promotes disassortative pollination between two floral morphs, has significant evolutionary implications. Here we examine the consequences of repeated loss of distyly in\n                  <jats:italic>Linum<\/jats:italic>\n                  by testing for relaxed selective pressure in homostylous relative to distylous lineages, and by characterising the evolutionary genomic patterns of the homostylous\n                  <jats:italic>Linum leonii<\/jats:italic>\n                  in comparison to its distylous close relative\n                  <jats:italic>Linum perenne<\/jats:italic>\n                  . We generated whole-genome sequences and target-capture data from sixteen\n                  <jats:italic>Linum<\/jats:italic>\n                  species, and additionally built a high-quality genome assembly and acquired population-level whole-genome sequencing data for\n                  <jats:italic>L. leonii<\/jats:italic>\n                  (n=20). We reconstructed plastome and nuclear phylogenies, estimated selective pressure for chloroplast and nuclear genes, inferred ancestral floral morph states, and tested for signatures of selfing in\n                  <jats:italic>L. leonii<\/jats:italic>\n                  . Compared to theoretical expectations, results were mixed, with partial identification of relaxed selective pressure in homostyles. One clade exhibited signs of potentially accelerated plastome evolution. Population genomic analyses of\n                  <jats:italic>L. leonii<\/jats:italic>\n                  revealed a moderate selfing rate of 0.32, suggesting that loss of distyly was associated with mixed mating rather than selfing, contrary to previous results on loss of distyly. Reduced nucleotide diversity and evidence for relaxed selection efficacy in\n                  <jats:italic>L. leonii<\/jats:italic>\n                  was likely due to a historical bottleneck. These results highlight the complex evolutionary dynamics associated with the breakdown of distyly. The genomic consequences are more heterogeneous than previously thought, and likely depend on clade- and species-specific evolutionary and demographic history dynamics. This study emphasizes the need for comparative population genomic studies to clarify how such transitions can shape evolutionary processes.\n                <\/jats:p>\n                <jats:sec>\n                  <jats:title>Significance statement<\/jats:title>\n                  <jats:p>\n                    Plant mating system variation is central to evolution as it shapes genetic diversity, adaptability and fitness. Loss of distyly, an iconic example of a complex mating system favouring cross-pollination, can drives shift from outcrossing to selfing, with potentially severe evolutionary consequences for the long-term persistence of the species in which it occurs. Using high-quality genome assembly and omic data for multiple\n                    <jats:italic>Linum<\/jats:italic>\n                    species, we tested for relaxed selective pressure following loss of distyly in homostylous species and tested for a signature of selfing in homostylous\n                    <jats:italic>Linum leonii<\/jats:italic>\n                    compared to the closely related distylous\n                    <jats:italic>Linum perenne<\/jats:italic>\n                    . Contrary to theoretical expectations, evidence for relaxation of selection was mixed in\n                    <jats:italic>Linum<\/jats:italic>\n                    homostyles and\n                    <jats:italic>L. leonii<\/jats:italic>\n                    did not exhibit a genomic signature of selfing. Our study reveals multiple evolutionary pathways following the loss of distyly, and highlights how mating system transitions, together with complex demographic processes, can differentially shape plant genetic diversity and evolution.\n                  <\/jats:p>\n                <\/jats:sec>","DOI":"10.64898\/2026.03.03.709227","type":"posted-content","created":{"date-parts":[[2026,3,5]],"date-time":"2026-03-05T00:25:15Z","timestamp":1772670315000},"source":"Crossref","is-referenced-by-count":0,"title":["Evolutionary consequences of repeated loss of distyly in\n                  <i>Linum<\/i>"],"prefix":"10.64898","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-0502-2375","authenticated-orcid":false,"given":"Zo\u00e9","family":"Postel","sequence":"first","affiliation":[{"name":"Department of Ecology, Environment and Plant Sciences, Science for Life Laboratory, Stockholm University, Stockholm, Sweden"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5197-3502","authenticated-orcid":false,"given":"Panagiotis-Ioannis","family":"Zervakis","sequence":"additional","affiliation":[{"name":"Department of Ecology, Environment and Plant Sciences, Science for Life Laboratory, Stockholm University, Stockholm, Sweden"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2962-2669","authenticated-orcid":false,"given":"Marco","family":"Fracassetti","sequence":"additional","affiliation":[{"name":"Department of Ecology, Environment and Plant Sciences, Science for Life Laboratory, Stockholm University, Stockholm, Sweden"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7669-9266","authenticated-orcid":false,"given":"Aleksandra","family":"Losvik","sequence":"additional","affiliation":[{"name":"Department of Ecology, Environment and Plant Sciences, Science for Life Laboratory, Stockholm University, Stockholm, Sweden"}]},{"given":"Matias","family":"Wanntorp","sequence":"additional","affiliation":[{"name":"Department of Ecology, Environment and Plant Sciences, Science for Life Laboratory, Stockholm University, Stockholm, Sweden"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-0121-2393","authenticated-orcid":false,"given":"Lucile","family":"Soler","sequence":"additional","affiliation":[{"name":"SciLifeLab, National Bioinformatics Infrastructure Sweden (NBIS), Department of Medical Biochemistry and Microbiology, Uppsala University, Uppsala, Sweden"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1902-3002","authenticated-orcid":false,"given":"Allison","family":"Churcher","sequence":"additional","affiliation":[{"name":"Department of Plant Physiology, National Bioinformatics Infrastructure Sweden, Science for Life Laboratory, Ume\u00e5 University, Ume\u00e5, Sweden"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2515-6509","authenticated-orcid":false,"given":"Aelys M.","family":"Humphreys","sequence":"additional","affiliation":[{"name":"Department of Ecology, Environment and Plant Sciences, Science for Life Laboratory, Stockholm University, Stockholm, Sweden"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-6020-5102","authenticated-orcid":false,"given":"Tanja","family":"Slotte","sequence":"additional","affiliation":[{"name":"Department of Ecology, Environment and Plant Sciences, Science for Life Laboratory, Stockholm University, Stockholm, Sweden"}]}],"member":"54368","reference":[{"key":"2026030703350562000_2026.03.03.709227v1.1","doi-asserted-by":"publisher","DOI":"10.1101\/gr.094052.109"},{"key":"2026030703350562000_2026.03.03.709227v1.2","doi-asserted-by":"publisher","DOI":"10.1534\/genetics.114.172809"},{"key":"2026030703350562000_2026.03.03.709227v1.3","doi-asserted-by":"publisher","DOI":"10.2307\/2405656"},{"key":"2026030703350562000_2026.03.03.709227v1.4","doi-asserted-by":"publisher","DOI":"10.1089\/cmb.2012.0021"},{"key":"2026030703350562000_2026.03.03.709227v1.5","doi-asserted-by":"publisher","DOI":"10.1186\/s12862-014-0268-y"},{"key":"2026030703350562000_2026.03.03.709227v1.6","doi-asserted-by":"publisher","DOI":"10.1111\/nph.16026"},{"key":"2026030703350562000_2026.03.03.709227v1.7","doi-asserted-by":"publisher","DOI":"10.1007\/978-3-540-68486-2_1"},{"key":"2026030703350562000_2026.03.03.709227v1.8","doi-asserted-by":"publisher","DOI":"10.1002\/1521-1878(200012)22:12<1075::AID-BIES5>3.0.CO;2-M"},{"key":"2026030703350562000_2026.03.03.709227v1.9","doi-asserted-by":"publisher","DOI":"10.1093\/sysbio\/syt034"},{"key":"2026030703350562000_2026.03.03.709227v1.10","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msaa042"},{"key":"2026030703350562000_2026.03.03.709227v1.11","doi-asserted-by":"publisher","DOI":"10.1111\/2041-210X.13534"},{"key":"2026030703350562000_2026.03.03.709227v1.12","doi-asserted-by":"publisher","DOI":"10.1016\/j.tig.2016.01.004"},{"key":"2026030703350562000_2026.03.03.709227v1.13","doi-asserted-by":"publisher","DOI":"10.1038\/nmeth.3176"},{"key":"2026030703350562000_2026.03.03.709227v1.14","doi-asserted-by":"publisher","DOI":"10.1002\/9780470015902.a0026804"},{"key":"2026030703350562000_2026.03.03.709227v1.15","unstructured":"Bushnell B. 2014. BBMap: A Fast, Accurate, Splice-Aware Aligner. Ernest Orlando Lawrence Berkeley National Laboratory, Berkeley, CA (US). Report No.: LBNL-7065E. [accessed 2025 July 31]. https:\/\/www.osti.gov\/biblio\/1241166"},{"key":"2026030703350562000_2026.03.03.709227v1.16","doi-asserted-by":"publisher","DOI":"10.1186\/1471-2105-10-421"},{"key":"2026030703350562000_2026.03.03.709227v1.17","doi-asserted-by":"publisher","DOI":"10.1186\/s13742-015-0047-8"},{"key":"2026030703350562000_2026.03.03.709227v1.18","doi-asserted-by":"publisher","DOI":"10.1086\/283497"},{"key":"2026030703350562000_2026.03.03.709227v1.19","doi-asserted-by":"publisher","DOI":"10.1086\/283496"},{"key":"2026030703350562000_2026.03.03.709227v1.20","doi-asserted-by":"publisher","DOI":"10.1016\/S0959-437X(00)00254-9"},{"key":"2026030703350562000_2026.03.03.709227v1.21","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msx088"},{"key":"2026030703350562000_2026.03.03.709227v1.22","doi-asserted-by":"publisher","DOI":"10.1038\/s41592-020-01056-5"},{"key":"2026030703350562000_2026.03.03.709227v1.23","doi-asserted-by":"publisher","DOI":"10.1111\/nph.16075"},{"key":"2026030703350562000_2026.03.03.709227v1.24","doi-asserted-by":"publisher","DOI":"10.1093\/bioinformatics\/btr330"},{"key":"2026030703350562000_2026.03.03.709227v1.25","doi-asserted-by":"publisher","DOI":"10.1017\/CBO9780511694202"},{"key":"2026030703350562000_2026.03.03.709227v1.26","doi-asserted-by":"crossref","unstructured":"Darwin C. 1877. The different forms of flowers on plants of the same species. The works of Charles Darwin (USA). [published online ahead of print] [accessed 2021 Mar 5]. https:\/\/agris.fao.org\/agris-search\/search.do?recordID=US9321022","DOI":"10.5962\/bhl.title.46988"},{"key":"2026030703350562000_2026.03.03.709227v1.27","doi-asserted-by":"publisher","DOI":"10.1017\/CBO9780511731419"},{"key":"2026030703350562000_2026.03.03.709227v1.28","doi-asserted-by":"publisher","DOI":"10.1126\/science.292.5517.673"},{"key":"2026030703350562000_2026.03.03.709227v1.29","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1412277112"},{"key":"2026030703350562000_2026.03.03.709227v1.30","doi-asserted-by":"publisher","DOI":"10.1111\/j.1558-5646.2010.01045.x"},{"key":"2026030703350562000_2026.03.03.709227v1.31","doi-asserted-by":"publisher","DOI":"10.1080\/0028825X.1979.10432574"},{"key":"2026030703350562000_2026.03.03.709227v1.32","doi-asserted-by":"publisher","DOI":"10.1534\/genetics.107.073601"},{"key":"2026030703350562000_2026.03.03.709227v1.33","doi-asserted-by":"publisher","DOI":"10.1111\/jeb.12356"},{"key":"2026030703350562000_2026.03.03.709227v1.34","doi-asserted-by":"publisher","DOI":"10.1146\/annurev.ecolsys.36.091704.175539"},{"key":"2026030703350562000_2026.03.03.709227v1.35","doi-asserted-by":"publisher","DOI":"10.1093\/nar\/gkz238"},{"key":"2026030703350562000_2026.03.03.709227v1.36","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pgen.1000695"},{"key":"2026030703350562000_2026.03.03.709227v1.37","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msae087"},{"key":"2026030703350562000_2026.03.03.709227v1.38","doi-asserted-by":"publisher","DOI":"10.1016\/j.tig.2017.04.002"},{"key":"2026030703350562000_2026.03.03.709227v1.39","doi-asserted-by":"publisher","DOI":"10.1016\/j.cub.2021.11.046"},{"key":"2026030703350562000_2026.03.03.709227v1.40","doi-asserted-by":"publisher","DOI":"10.1111\/nph.12182"},{"key":"2026030703350562000_2026.03.03.709227v1.41","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.0709121104"},{"key":"2026030703350562000_2026.03.03.709227v1.42","doi-asserted-by":"publisher","DOI":"10.1111\/mec.15684"},{"key":"2026030703350562000_2026.03.03.709227v1.43","doi-asserted-by":"publisher","DOI":"10.1186\/s13059-020-02154-5"},{"key":"2026030703350562000_2026.03.03.709227v1.44","doi-asserted-by":"publisher","DOI":"10.3732\/apps.1600016"},{"key":"2026030703350562000_2026.03.03.709227v1.45","doi-asserted-by":"publisher","DOI":"10.1093\/sysbio\/syy086"},{"key":"2026030703350562000_2026.03.03.709227v1.46","doi-asserted-by":"publisher","DOI":"10.1080\/17550874.2021.2012849"},{"key":"2026030703350562000_2026.03.03.709227v1.47","doi-asserted-by":"publisher","DOI":"10.1534\/genetics.118.301120"},{"key":"2026030703350562000_2026.03.03.709227v1.48","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1707492115"},{"key":"2026030703350562000_2026.03.03.709227v1.49","doi-asserted-by":"publisher","DOI":"10.1093\/aob\/mcr159"},{"key":"2026030703350562000_2026.03.03.709227v1.50","doi-asserted-by":"publisher","DOI":"10.48550\/arXiv.1303.3997"},{"key":"2026030703350562000_2026.03.03.709227v1.51","doi-asserted-by":"publisher","DOI":"10.1093\/bioinformatics\/btp698"},{"key":"2026030703350562000_2026.03.03.709227v1.52","doi-asserted-by":"publisher","DOI":"10.1007\/978-3-642-86656-2_6"},{"key":"2026030703350562000_2026.03.03.709227v1.53","doi-asserted-by":"publisher","DOI":"10.1111\/jbi.14129"},{"key":"2026030703350562000_2026.03.03.709227v1.54","doi-asserted-by":"publisher","DOI":"10.1007\/978-1-0716-0199-0_11"},{"key":"2026030703350562000_2026.03.03.709227v1.55","doi-asserted-by":"publisher","DOI":"10.1600\/036364409788606244"},{"key":"2026030703350562000_2026.03.03.709227v1.56","doi-asserted-by":"publisher","DOI":"10.1111\/mec.16270"},{"key":"2026030703350562000_2026.03.03.709227v1.57","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msaa229"},{"key":"2026030703350562000_2026.03.03.709227v1.58","doi-asserted-by":"publisher","DOI":"10.1006\/jmbi.2000.4042"},{"issue":"4","key":"2026030703350562000_2026.03.03.709227v1.59","doi-asserted-by":"crossref","first-page":"787","DOI":"10.1111\/j.1469-8137.1968.tb06396.x","article-title":"Biosystematic Studies in the Linum perenne Group","volume":"67","year":"1968","journal-title":"The New Phytologist"},{"key":"2026030703350562000_2026.03.03.709227v1.60","doi-asserted-by":"publisher","DOI":"10.1186\/s13059-016-0997-x"},{"key":"2026030703350562000_2026.03.03.709227v1.61","doi-asserted-by":"publisher","DOI":"10.1126\/science.1180677"},{"key":"2026030703350562000_2026.03.03.709227v1.62","doi-asserted-by":"publisher","DOI":"10.1093\/bioinformatics\/bty633"},{"key":"2026030703350562000_2026.03.03.709227v1.63","doi-asserted-by":"publisher","DOI":"10.1111\/mec.12524"},{"key":"2026030703350562000_2026.03.03.709227v1.64","doi-asserted-by":"publisher","DOI":"10.1016\/j.ympev.2022.107436"},{"key":"2026030703350562000_2026.03.03.709227v1.65","doi-asserted-by":"publisher","DOI":"10.7717\/peerj.16505"},{"key":"2026030703350562000_2026.03.03.709227v1.66","doi-asserted-by":"publisher","DOI":"10.1534\/genetics.116.188268"},{"key":"2026030703350562000_2026.03.03.709227v1.67","doi-asserted-by":"publisher","DOI":"10.1016\/bs.abr.2017.11.017"},{"key":"2026030703350562000_2026.03.03.709227v1.68","doi-asserted-by":"publisher","DOI":"10.1111\/plb.12670"},{"key":"2026030703350562000_2026.03.03.709227v1.69","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msae070"},{"key":"2026030703350562000_2026.03.03.709227v1.70","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0163962"},{"key":"2026030703350562000_2026.03.03.709227v1.71","doi-asserted-by":"publisher","DOI":"10.1093\/gbe\/evaa123"},{"key":"2026030703350562000_2026.03.03.709227v1.72","doi-asserted-by":"publisher","DOI":"10.1093\/aob\/mcr023"},{"key":"2026030703350562000_2026.03.03.709227v1.73","doi-asserted-by":"publisher","DOI":"10.1038\/s41467-024-45118-0"},{"key":"2026030703350562000_2026.03.03.709227v1.74","doi-asserted-by":"publisher","DOI":"10.1016\/j.ympev.2013.12.004"},{"key":"2026030703350562000_2026.03.03.709227v1.75","doi-asserted-by":"publisher","DOI":"10.1093\/bfgp\/elu009"},{"key":"2026030703350562000_2026.03.03.709227v1.76","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msq062"},{"key":"2026030703350562000_2026.03.03.709227v1.77","doi-asserted-by":"publisher","DOI":"10.1093\/bioinformatics\/btu033"},{"key":"2026030703350562000_2026.03.03.709227v1.78","doi-asserted-by":"publisher","DOI":"10.1093\/aob\/mct306"},{"key":"2026030703350562000_2026.03.03.709227v1.79","doi-asserted-by":"publisher","DOI":"10.2307\/3558325"},{"key":"2026030703350562000_2026.03.03.709227v1.80","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msab120"},{"key":"2026030703350562000_2026.03.03.709227v1.81","doi-asserted-by":"publisher","DOI":"10.1093\/nar\/gkx391"},{"key":"2026030703350562000_2026.03.03.709227v1.82","doi-asserted-by":"publisher","DOI":"10.3390\/agronomy14010099"},{"key":"2026030703350562000_2026.03.03.709227v1.83","doi-asserted-by":"publisher","DOI":"10.1098\/rstb.2020.0510"},{"key":"2026030703350562000_2026.03.03.709227v1.84","doi-asserted-by":"publisher","DOI":"10.1086\/695527"},{"key":"2026030703350562000_2026.03.03.709227v1.85","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msaa199"},{"key":"2026030703350562000_2026.03.03.709227v1.86","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msx319"},{"key":"2026030703350562000_2026.03.03.709227v1.87","doi-asserted-by":"publisher","DOI":"10.1016\/j.ympev.2012.05.026"},{"key":"2026030703350562000_2026.03.03.709227v1.88","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1323926111"},{"key":"2026030703350562000_2026.03.03.709227v1.89","doi-asserted-by":"publisher","DOI":"10.1007\/BF02459575"},{"key":"2026030703350562000_2026.03.03.709227v1.90","doi-asserted-by":"publisher","DOI":"10.1111\/j.1469-1809.1949.tb02451.x"},{"key":"2026030703350562000_2026.03.03.709227v1.91","unstructured":"Wright S . 1984. Evolution and the Genetics of Populations, Volume 2: Theory of Gene Frequencies. University of Chicago Press."},{"key":"2026030703350562000_2026.03.03.709227v1.92","doi-asserted-by":"publisher","DOI":"10.1098\/rspb.2013.0133"},{"key":"2026030703350562000_2026.03.03.709227v1.93","doi-asserted-by":"publisher","DOI":"10.1086\/523366"},{"key":"2026030703350562000_2026.03.03.709227v1.94","doi-asserted-by":"publisher","DOI":"10.1093\/molbev\/msm088"},{"key":"2026030703350562000_2026.03.03.709227v1.95","doi-asserted-by":"publisher","DOI":"10.1093\/aob\/mcx098"},{"key":"2026030703350562000_2026.03.03.709227v1.96","doi-asserted-by":"publisher","DOI":"10.1038\/s41437-018-0081-2"},{"key":"2026030703350562000_2026.03.03.709227v1.97","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.2214492120"},{"key":"2026030703350562000_2026.03.03.709227v1.98","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pgen.1010883"},{"key":"2026030703350562000_2026.03.03.709227v1.99","doi-asserted-by":"publisher","DOI":"10.1111\/nph.70392"},{"key":"2026030703350562000_2026.03.03.709227v1.100","doi-asserted-by":"publisher","DOI":"10.1186\/s12859-018-2129-y"},{"key":"2026030703350562000_2026.03.03.709227v1.101","doi-asserted-by":"publisher","DOI":"10.1105\/tpc.114.134353"},{"key":"2026030703350562000_2026.03.03.709227v1.102","doi-asserted-by":"publisher","DOI":"10.1111\/nph.15905"}],"container-title":[],"original-title":[],"link":[{"URL":"https:\/\/syndication.highwire.org\/content\/doi\/10.64898\/2026.03.03.709227","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,3,7]],"date-time":"2026-03-07T11:35:17Z","timestamp":1772883317000},"score":1,"resource":{"primary":{"URL":"http:\/\/biorxiv.org\/lookup\/doi\/10.64898\/2026.03.03.709227"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,3,4]]},"references-count":102,"URL":"https:\/\/doi.org\/10.64898\/2026.03.03.709227","relation":{},"subject":[],"published":{"date-parts":[[2026,3,4]]},"subtype":"preprint"}}