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The species is compared with its congener B. hugoi Spinelli &amp; Grogan, and a key to females and males of the six Neotropical species of the subgenus is included<\/jats:p>","DOI":"10.11646\/zootaxa.570.1.1","type":"journal-article","created":{"date-parts":[[2016,9,26]],"date-time":"2016-09-26T01:54:25Z","timestamp":1474854865000},"source":"Crossref","is-referenced-by-count":0,"title":["A new species of Brachypogon (Isohelea) from arid zones of Argentina (Diptera: Ceratopogonidae)"],"prefix":"10.11646","volume":"570","author":[{"given":"GUSTAVO R.","family":"SPINELLI","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"CARLA G.","family":"CAZORLA","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2004,7,16]]},"reference":[{"key":"27512","unstructured":"<p>Borkent, A. &amp; Wirth, W.W. (1997) World species of biting midges (Diptera: Ceratopogonidae). <em>Bulletin of the American Museum of Natural History<\/em>, 233, 1\u2013257.<\/p><p>Borkent, A. &amp; Spinelli, G.R. (2000) Catalog of the New World biting midges south of the United States of America (Diptera: Ceratopogonidae). <em>Contributions on Entomology<\/em>, <em>International<\/em>, 4, 1\u2013107.<\/p><p>McAlpine, J.F., Peterson, B.V, Shewell, G.E, Teskey, H.J, Vockeroth, J.R. &amp; Wood, D.M. (coordinators) (1981) <em>Manual of Nearctic Diptera<\/em>. Volume 1. Agriculture Canada Monograph, 27, 1\u2013 674.<\/p><p>Spinelli, G.R. &amp; Grogan, W.L. (1994) Two new Neotropical species of the subgenus <em>Isohelea <\/em>of <em>Brachypogon<\/em>, with the description of the female of <em>B<\/em>. (<em>I<\/em>.) <em>misionensis <\/em>and a key to the Neotropical species (Diptera: Ceratopogonidae). <em>Revista de la Sociedad Entomol\u00f3gica Argentina<\/em>, 53, 1\u20138.<\/p><p>Spinelli, G.R. &amp; Grogan, W.L. (1998) A revision of the Neotropical predaceous midges of <em>Brachypogon <\/em>(<em>Brachypogon<\/em>) Kieffer (Diptera: Ceratopogonidae). <em>Insecta Mundi<\/em>, 12, 59\u201379.<\/p><p>Szadziewski, R. (1996) Biting midges from Lower Cretaceous amber of Lebanon and Upper Cretaceous Siberian amber of Taimyr (Diptera, Ceratopogonidae). <em>Studia Dipterologica<\/em>, 3, 23\u201386.<\/p><p>Szadziewski, R. &amp; Grogan, W.L. (1998) Biting midges from Dominican amber. III. Species of the tribes Culicoidini and Ceratopogonini (Diptera: Ceratopogonidae). <em>Insecta Mundi<\/em>, 12, 39\u201352.<\/p>"}],"container-title":["Zootaxa"],"link":[{"URL":"https:\/\/mapress.com\/zt\/article\/download\/zootaxa.570.1.1\/8042","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/mapress.com\/zt\/article\/download\/zootaxa.570.1.1\/8042","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,3,19]],"date-time":"2024-03-19T00:20:01Z","timestamp":1710807601000},"score":14.561722,"resource":{"primary":{"URL":"https:\/\/mapress.com\/zt\/article\/view\/zootaxa.570.1.1"}},"issued":{"date-parts":[[2004,7,16]]},"references-count":1,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2004,7,16]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.570.1.1","ISSN":["1175-5334","1175-5326"],"issn-type":[{"value":"1175-5334","type":"electronic"},{"value":"1175-5326","type":"print"}],"published":{"date-parts":[[2004,7,16]]}},{"indexed":{"date-parts":[[2025,2,21]],"date-time":"2025-02-21T20:34:57Z","timestamp":1740170097169,"version":"3.37.3"},"reference-count":1,"publisher":"Magnolia Press","issue":"4","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"<jats:p>The predaceous midge genus Brachypogon Kieffer is presently known in Mexico by Brachypogon (Brachypogon) fuscivenosus (Lutz, 1914), B. (B.) bimaculatus Spinelli &amp; Grogan, 1998, B. (B.) woodruffi Spinelli &amp; Grogan, 1998 and B. (Isohelea) cuacuahuitlus Huerta &amp; Borkent, 2005. In this contribution a new species, Brachypogon (Brachypogon) ginue, is described and illustrated from a male specimen collected in Tlanchinol, State of Hidalgo, Mexico. This new species belongs to the impar species group. The first description of the female of Brachypogon (Isohelea) cuacuahuitlus Huerta &amp; Borkent is also provided.\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.4066.4.8","type":"journal-article","created":{"date-parts":[[2016,1,17]],"date-time":"2016-01-17T23:58:42Z","timestamp":1453075122000},"source":"Crossref","is-referenced-by-count":0,"title":["A new species of the predaceous midge genus Brachypogon Kieffer from the Neotropical Region and first description of the female of Brachypogon (Isohelea) cuacuahuitlus Huerta &amp;amp; Borkent (Diptera: Ceratopogonidae)"],"prefix":"10.11646","volume":"4066","author":[{"given":"HER\u00d3N","family":"HUERTA","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"GUSTAVO R.","family":"SPINELLI","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2016,1,18]]},"reference":[{"key":"37246","unstructured":"<p class=\"HeadingRunIn\">Borkent, A. (2015) World species of biting midges (Diptera: Ceratopogonidae): On line catalog. FLYTREE Assembling the Diptera Tree of Life, Art Borkent. Available from: h<a href=\"http:\/\/wwx.inhs.illinois.edu\/files\/1114\/2384\/5200\/CeratopogonidaeCatalog.pdf\">ttp:\/\/wwx.inhs.illinois.edu\/files\/1114\/2384\/5200\/<\/a>CeratopogonidaeCatalog.pdf (accessed 7 October 2015)<\/p><p class=\"HeadingRunIn\">Borkent, A. &amp; Grogan, W.L.Jr. (2009) Catalog of the New World biting midges North of Mexico (Ceratopogonidae: Diptera). <em>Zootaxa<\/em>, 2273, 1\u201348.<\/p><p class=\"HeadingRunIn\">Borkent, A. &amp; Spinelli, G.R. (2007) Neotropical Ceratopogonidae (Diptera: Insecta). <em>In<\/em>: Adis, J., Arias, J.R., Rueda-Delgado, G. &amp; Wantzen, K.M. (Eds.), <em>Aquatic Biodiversity in Latin America (ABLA). Vol. 4.<\/em> Pensoft, Sofia-Moscow, pp. 1\u2013198.<\/p><p class=\"HeadingRunIn\">Borkent, A., Spinelli, G.R. &amp; Grogan, W.L. Jr. (2009) Ceratopogonidae (Biting midges, purrujas). <em>In<\/em>: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. &amp; Zumbado, M.A. (Eds.), <em>Manual of Central American Diptera<\/em>. <em>Vol. 1<\/em>. National Research Council Research Press, Ottawa, Ontario, pp. 407\u2013435.<\/p><p class=\"HeadingRunIn\">Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. &amp; Zumbado, M.A. (2009) <em>Manual of Central American Diptera<\/em>. <em>Vol. 1.<\/em> National Research Council Research Press, Ottawa, 714 pp.<\/p><p class=\"HeadingRunIn\">Grogan, W.L.Jr., Spinelli, G.R., Ronderos, M.M. &amp; Cazorla, C.G. (2013) The biting and predaceous midges of Guadeloupe (Diptera: Ceratopogonidae). I. Species of the subfamily Ceratopogoninae. <em>Insecta Mundi<\/em>, 0324, 1\u201321.<\/p><p class=\"HeadingRunIn\">Huerta, H. &amp; Borkent, A. (2005) A new species and first record of <em>Ceratoculicoides<\/em> With and Ratanaworabhan from the Neotropical Region and new species and records of <em>Brachypogon <\/em>Kieffer from Mexico (Diptera: Ceratopogonidae). <em>Folia Entomol\u00f3gica Mexicana,<\/em> 44 (Supplement 1), 111\u2013119.<\/p><p class=\"HeadingRunIn\"> <\/p><p class=\"HeadingRunIn\">Lutz, A. (1914) Contribui\u00e7\u00e3o para o conhecimento das \u201cCeratopogoninas\u201d do Brasil. <em>Memorias do Instituto Oswaldo Cruz,<\/em> 6, 81\u201399.<\/p><p class=\"HeadingRunIn\">Spinelli, G.R. (1990) The genus <em>Brachypogon<\/em> in Argentina, with a key to the Neotropical species (Diptera: Ceratopogonidae). <em>Revista Brasileira de Entomologia<\/em>, 34 (4), 743\u2013755.<\/p><p class=\"HeadingRunIn\">Spinelli, G.R., Borkent, A. &amp; Ronderos, M.M. (2013) Two new peculiar species of Neotropical <em>Brachypogon <\/em>Kieffer (Diptera: Ceratopogonidae). <em>Zootaxa<\/em>, 3702 (1), 90\u201396.<br \/> http:\/\/dx.doi.org\/10.11646\/zootaxa.3702.1.6<\/p><p class=\"HeadingRunIn\">Spinelli, G.R. &amp; Cazorla, C.G. (2004) A new species of <em>Brachypogon <\/em>(<em>Isohelea<\/em>) from arid zones of Argentina (Diptera: Ceratopogonidae). <em>Zootaxa<\/em>, 570, 1\u20136.<\/p><p class=\"HeadingRunIn\">Spinelli, G.R. &amp; Grogan, W.L. Jr. (1994) Two new Neotropical species of the subgenus <em>Isohelea <\/em>of <em>Brachypogon<\/em>, with the description of the female of <em>B. (I.) misionensis<\/em> and a key to the Neotropical species (Diptera: Ceratopogonidae). <em>Revista de la Sociedad Entomol\u00f3gica Argentina<\/em>, 53 (1\u20134), 1\u20138.<\/p><p class=\"HeadingRunIn\">Spinelli, G.R. &amp; Grogan, W.L. Jr. (1998) A revision of the Neotropical predaceous midges of <em>Brachypogon (Brachypogon<\/em>) Kieffer (Diptera: Ceratopogonidae). <em>Insecta Mundi<\/em>, 12 (1\u20132), 59\u201379.<\/p><p class=\"HeadingRunIn\">Swanson, D.A. &amp; Grogan, W.L. Jr. (2011) A new predaceous midge in <em>Brachypogon <\/em>(<em>Brachypogon<\/em>) from Alabama and Florida, U.S.A. and new distribution records for <em>Brachypogon woodruffi <\/em>Spinelli and Grogan (Diptera: Ceratopogonidae). <em>Proceedings of the Entomological Society of Washington<\/em>, 113, 531\u2013540.<\/p><p> <\/p>"}],"container-title":["Zootaxa"],"link":[{"URL":"https:\/\/mapress.com\/zt\/article\/download\/zootaxa.4066.4.8\/3825","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/mapress.com\/zt\/article\/download\/zootaxa.4066.4.8\/3825","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,3,18]],"date-time":"2024-03-18T00:10:33Z","timestamp":1710720633000},"score":14.0933895,"resource":{"primary":{"URL":"https:\/\/mapress.com\/zt\/article\/view\/zootaxa.4066.4.8"}},"issued":{"date-parts":[[2016,1,18]]},"references-count":1,"journal-issue":{"issue":"4","published-online":{"date-parts":[[2016,1,18]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.4066.4.8","ISSN":["1175-5334","1175-5326"],"issn-type":[{"type":"electronic","value":"1175-5334"},{"type":"print","value":"1175-5326"}],"published":{"date-parts":[[2016,1,18]]}},{"indexed":{"date-parts":[[2024,4,16]],"date-time":"2024-04-16T23:22:11Z","timestamp":1713309731787},"reference-count":0,"publisher":"Societe entomologique de France","issue":"5","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["bsef"],"published-print":{"date-parts":[[1989]]},"DOI":"10.3406\/bsef.1989.17601","type":"journal-article","created":{"date-parts":[[2021,11,9]],"date-time":"2021-11-09T09:07:15Z","timestamp":1636448835000},"page":"189-196","source":"Crossref","is-referenced-by-count":1,"title":["Description de trois Brachypogon nouveaux et remarquables de la R\u00e9publique de Guin\u00e9e [Dipt. Ceratopogonidae]"],"prefix":"10.32475","volume":"94","author":[{"given":"Jean","family":"Clastrier","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"17575","container-title":["Bulletin de la Soci\u00e9t\u00e9 entomologique de France"],"language":"fr","deposited":{"date-parts":[[2021,11,13]],"date-time":"2021-11-13T05:52:18Z","timestamp":1636782738000},"score":14.058819,"resource":{"primary":{"URL":"https:\/\/www.persee.fr\/doc\/bsef_0037-928x_1989_num_94_5_17601"}},"issued":{"date-parts":[[1989]]},"references-count":0,"journal-issue":{"issue":"5","published-print":{"date-parts":[[1989]]}},"URL":"https:\/\/doi.org\/10.3406\/bsef.1989.17601","ISSN":["0037-928X"],"issn-type":[{"value":"0037-928X","type":"print"}],"published":{"date-parts":[[1989]]}},{"indexed":{"date-parts":[[2025,5,14]],"date-time":"2025-05-14T03:45:24Z","timestamp":1747194324832,"version":"3.40.5"},"reference-count":18,"publisher":"Magnolia Press","issue":"1","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"<jats:p>N<\/jats:p>","DOI":"10.11646\/zootaxa.5319.1.12","type":"journal-article","created":{"date-parts":[[2023,7,24]],"date-time":"2023-07-24T00:20:45Z","timestamp":1690158045000},"page":"145-147","source":"Crossref","is-referenced-by-count":0,"title":["Taxonomic status of two European sibling and barcode-sharing species of Brachypogon Kieffer, 1899 (Diptera: Ceratopogonidae)"],"prefix":"10.11646","volume":"5319","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-0034-069X","authenticated-orcid":false,"given":"PATRYCJA","family":"DOMINIAK","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2899-6861","authenticated-orcid":false,"given":"RYSZARD","family":"SZADZIEWSKI","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2023,7,24]]},"reference":[{"key":"218327","doi-asserted-by":"crossref","unstructured":"Baselga, A., G\u00f3mez-Rodr\u00edguez, C., Novoa, F. & Vogler, A.P. (2013) Rare failures of DNA bar codes to separate morphologically distinct species in a biodiversity survey of Iberian leaf beetles. PLoS ONE, 8 (9), e74854. https:\/\/doi.org\/10.1371\/annotation\/18a13c3f-ea86-45bf-9934-35d9bde2029c","DOI":"10.1371\/journal.pone.0074854"},{"key":"218328","doi-asserted-by":"crossref","unstructured":"Borkent, A. & Dominiak, P. (2020) Catalog of the biting midges of the world (Diptera: Ceratopogonidae). Zootaxa, 4787 (1), 1\u2013377. https:\/\/doi.org\/10.11646\/zootaxa.4787.1.1","DOI":"10.11646\/zootaxa.4787.1.1"},{"key":"218329","doi-asserted-by":"crossref","unstructured":"Borkent, A., Dominiak, P. & D\u00edaz, F. (2022) An update and errata for the catalog of the biting midges of the world (Diptera: Ceratopogonidae). Zootaxa, 5120 (1), 53\u201364. https:\/\/doi.org\/10.11646\/zootaxa.5120.1.3","DOI":"10.11646\/zootaxa.5120.1.3"},{"key":"218330","unstructured":"Clastrier, J. (1961) Notes sur les C\u00e9ratopogonid\u00e9s. XV.-Ceratopogon et Alluaudomyia de la R\u00e9gion Pal\u00e9arctique. Archives de l\u2019Institut Pasteur d\u2019Alg\u00e9rie, 39, 401\u2013437."},{"key":"218331","unstructured":"Del\u00e9colle, J.-C. & Grogan, W.L. (1990) Two new species of the predaceous midge genus Brachypogon from S\u00e9n\u00e9gal (Diptera, Ceratopogonidae). Nouvelle Revue d'Entomologie, 7, 127\u2013138."},{"key":"218332","unstructured":"Dominiak, P. & Stur, E. (2022) New findings and an overall assessment of Norwegian biting midges. Norwegian Journal of Entomology, 69, 82\u2013190."},{"key":"218333","doi-asserted-by":"crossref","unstructured":"Kieffer, J.J. (1899) Description d\u2019un nouveau genre et tableau des genres europ\u00e9ens de la famille des Chironomides (Dipt.). Bulletin de la Soci\u00e9t\u00e9 Entomologique de France, 1899, 66\u201370. https:\/\/doi.org\/10.5962\/bhl.part.8618","DOI":"10.5962\/bhl.part.8618"},{"key":"218334","unstructured":"Kieffer, J.J. (1925) Nouveaux genres et nouvelles esp\u00e8ces de chironomides piqueurs. Archives de l\u2019Institut Pasteur d\u2019Alg\u00e9rie, 3, 405\u2013430."},{"key":"218335","doi-asserted-by":"crossref","unstructured":"Kj\u00e6randsen, J. (2022) Current state of DNA barcoding of Sciaroidea (Diptera)\u2014highlighting the need to build the reference library. Insects, 13 (2), 147. https:\/\/doi.org\/10.3390\/insects13020147","DOI":"10.3390\/insects13020147"},{"key":"218336","doi-asserted-by":"crossref","unstructured":"Kurina, O., \u00d5unap, E. & P\u00f5ldmaa, K. (2015) Two new Neuratelia Rondani (Diptera, Mycetophilidae) species from Western Palaearctic: a case of limited congruence between morphology and DNA sequence data. ZooKeys, 496, 105\u2013129. https:\/\/doi.org\/10.3897\/zookeys.496.9315","DOI":"10.3897\/zookeys.496.9315"},{"key":"218337","unstructured":"Edwards, F.W. (1921) Diptera Nematocera from Arran and Loch Etive. The Scottish Naturalist, 1921, 59\u201361 + 89\u201392 + 121\u2013125."},{"key":"218338","doi-asserted-by":"crossref","unstructured":"Edwards, F.W. (1926) On the British biting midges (Diptera, Ceratopogonidae). Transactions of the Entomological Society of London, 74, 389\u2013426, 2 pls. https:\/\/doi.org\/10.1111\/j.1365-2311.1926.tb02249.x","DOI":"10.1111\/j.1365-2311.1926.tb02249.x"},{"key":"218339","unstructured":"Remm, H. (1974) A systematic review of species of the genus Ceratopogon Meigen (Diptera) from USSR. Tartu Riikliku Ulikooli Toimetised, 327, 23\u201358. [in Russian, English summary]"},{"key":"218340","unstructured":"Remm, H. (1981) New synonyms and new names of the Palaearctic Ceratopogonidae (Diptera). Eesti NSV Teaduste Akadeemia Toimetised, 30, 27\u201332."},{"key":"218341","unstructured":"Remm, H. (1988) Family Ceratopogonidae. In: Soos, A. (Ed.), Catalogue of Palaearctic Diptera. Vol. 3. Ceratopogonidae-Mycetophilidae. Akad\u00e9miai Kiad\u00f3, Budapest, pp. 11\u2013110."},{"key":"218342","doi-asserted-by":"crossref","unstructured":"Stur, E. & Borkent, A. (2014) When DNA barcoding and morphology mesh: Ceratopogonidae diversity in Finnmark, Norway. ZooKeys, 463, 95\u2013131. https:\/\/doi.org\/10.3897\/zookeys.463.7964","DOI":"10.3897\/zookeys.463.7964"},{"key":"218343","unstructured":"Szadziewski, R. & Hagan, D.V. (2000) Two new species of biting midges from Norway (Diptera: Ceratopogonidae). Polskie Pismo Entomologiczne, 69, 459\u2013464."},{"key":"218344","unstructured":"Szadziewski, R., Kaczorowska, E. & Krzywi\u0144ski, J. (1994) The predaceous midges of the subgenus Isohelea of Brachypogon in Poland (Diptera, Ceratopogonidae). Acta Zoologica Cracoviensia, 37, 1\u201332."}],"container-title":["Zootaxa"],"deposited":{"date-parts":[[2023,7,24]],"date-time":"2023-07-24T00:20:57Z","timestamp":1690158057000},"score":13.256578,"resource":{"primary":{"URL":"https:\/\/mapress.com\/zt\/article\/view\/zootaxa.5319.1.12"}},"issued":{"date-parts":[[2023,7,24]]},"references-count":18,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2023,7,24]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.5319.1.12","ISSN":["1175-5334","1175-5326"],"issn-type":[{"type":"electronic","value":"1175-5334"},{"type":"print","value":"1175-5326"}],"published":{"date-parts":[[2023,7,24]]}},{"indexed":{"date-parts":[[2025,11,27]],"date-time":"2025-11-27T01:11:30Z","timestamp":1764205890424,"version":"3.46.0"},"reference-count":0,"publisher":"CSIRO Publishing","issue":"4","license":[{"start":{"date-parts":[[2025,4,22]],"date-time":"2025-04-22T00:00:00Z","timestamp":1745280000000},"content-version":"vor","delay-in-days":12318,"URL":"https:\/\/doi.org\/10.1071\/journalslicense"}],"content-domain":{"domain":[],"crossmark-restriction":false},"published-print":{"date-parts":[[1991,8,1]]},"abstract":"<jats:p>The status and differentiation of the genera Ceratopogon, Brachypogon and Isohelea have remained uncertain despite repeated attempts at clarification. In 1964 Tokunaga described five species of Ceratopogon from Papua New Guinea, treating Brachypogon and Isohelea as synonyms of Ceratopogon. Wirth &amp; Grogan (1988) determined that Ceratopogon was a purely Holarctic genus, and listed four of Tokunaga's species in Brachypogon, which they regarded as a genus with two subgenera, Brachypogon s.s. and Isohelea. (The fifth species was transferred to another genus, Nannohelea.) No species of the Ceratopogon group have been described from Australia. In this paper 24 species, 20 of which are new, are recorded from Australia and New Guinea. Seventeen of the new species \u2014 B. artemis, B. bryanae, B. corniger, B. medusae, B. bifurcus, B. pollices, B. tabernaculum, B. gearyae, B. hercules, B. subiectus, B. gravidus, B. idolon, B. lorica, B. curtus, B. institor, B. griffithsae and B. nicolaii \u2014 and two of Tokunaga's species, B. petersi and B. novaguineae, are placed in Brachypogon (Brachypogon), in two species-groups. Two of the new species, B. hadrosaurus and B. dehiscens, are placed in Brachypogon (Isohelea), while B. alexandros, sp. nov. and Tokunaga's B. maai and B. papuensis are placed in a new subgenus of Brachypogon, Sarissohelea. 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