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Drilling, K., Dettner, K. & Klass, K.D. (2013) The distribution and evolution of exocrine compound glands in Erotylinae (Insecta: Coleoptera: Erotylidae). Annales de la Soci\u00e9t\u00e9 Entomologique de France<\/em>, 49 (1), 36\u201352.<\/p>

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Lawrence, J.F., \u015alipi\u0144ski, A., Seago, A.E., Thayer, M.K., Newton, A.F. & Marvaldi, A.E. (2011) Phylogeny of the Coleoptera based on morphological characters of adults and larvae. Annales Zoologici, Warszawa<\/em>, 61, 1\u2013217.<\/p>

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Leschen, R.A.B. (2003) Erotylidae (Insecta: Coleoptera: Cucujoidea): phylogeny and review. Fauna of New Zealand<\/em>, 47, 1\u2013108.<\/p>

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Leschen, R.A.B. & W\u0119grzynowicz, P. (2008) Arrowcryptus<\/em>, a new genus of Xenoscelinae of Erotylidae from Africa. Zootaxa<\/em>, 1968 (1), 45\u201357.<\/p>

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Lyubarsky, G.Y. & Perkovsky, E.E. (2012) A new genus of Erotylidae from Eocene amber (Coleoptera: Clavicornia). Russian Entomological Journal<\/em>, 21 (1), 35\u201338.<\/p>

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Lyubarsky, G.Y. & Perkovsky, E.E. (2017a) Serramorphus<\/em>, a new genus of Erotylidae from Eocene amber (Coleoptera: Clavicornia) from Late Eocene Bitterfeldian amber. Russian Entomological Journal<\/em>, 26 (1), 37\u201340.<\/p>

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Lyubarsky, G.Y. & Perkovsky, E.E. (2017b) Xenophagus<\/em>, a new genus of pleasing fungus beetles (Coleoptera: Erotylidae) from Baltic amber. Russian Entomological Journal<\/em>, 26 (2), 147\u2013150.<\/p>

https:\/\/doi.org\/10.15298\/rusentj.26.2.05<\/a><\/p>

Lyubarsky, G.Y. & Perkovsky, E.E. (2018) Microzavaljus<\/em>, a second extinct genus of pleasing fungus beetles (Coleoptera: Erotylidae) from Bitterfeld Amber. Russian Entomological Journal<\/em>, 27 (3), 271\u2013276.<\/p>

https:\/\/doi.org\/10.15298\/rusentj.27.3.05<\/a><\/p>

Lyubarsky, G.Y., Perkovsky, E.E. & Alekseev, V.I. (2016) The first record of the subfamily Xenoscelinae (Coleoptera, Erotylidae) from the Baltic amber. Paleontological Journal<\/em>, 50 (9), 963\u2013969.<\/p>

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Robertson, J.A., McHugh, J.V. & Whiting, M.F. (2004) A molecular phylogenetic analysis of the pleasing fungus beetles (Coleoptera: Erotylidae): evolution of colour patterns, gregariousness and mycophagy. Systematic Entomology<\/em>, 29 (2), 173\u2013187.<\/p>

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Sen Gupta, T. & Crowson, R.A. (1971) A review of classification of the family Languriidae (Coleoptera: Clavicornia) and the place of Languriidae in the natural system of Clavicornia. Memoires of the Zoological Survey of India<\/em>, 15, 1\u201342.<\/p>

Skelley, P.E. & Powell, G.S. (2018) Necessary nomenclatural corrections for Mycophtorus<\/em> Lacordaire, 1842 and Neomycotretus<\/em> Deelder, 1942, with the establishment of Myceporthus<\/em> Skelley and Powell, new genus (Coleoptera: Erotylidae: Tritomini). The Coleopterists Bulletin<\/em>, 72 (2), 305\u2013313.<\/p>

https:\/\/doi.org\/10.1649\/0010-065X-72.2.305<\/a><\/p>

Skelley, P., Xu, G., Tang, W., Lindstr\u00f6m, A.J., Marler, T., Khuraijam, J.S., Singh, R., Radha, P. & Rich, S. (2017) Review of Cycadophila<\/em> Xu, Tang & Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas<\/em> (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species. Zootaxa<\/em>, 4267 (1), 1\u201363.<\/p>

https:\/\/doi.org\/10.11646\/zootaxa.4267.1.1<\/a><\/p>

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https:\/\/doi.org\/10.11646\/zootaxa.3148.1.39<\/a><\/p>

Tang, W., Skelley, P. & P\u00e9rez-Farrera, M.A. (2018) Ceratophila<\/em>, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia<\/em> (Cycadales: Zamiaceae). Zootaxa<\/em>, 4508 (2), 151\u2013178.<\/p>

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Xu, G., Tang, W., Skelley, P., Liu, N. & Rich, S. (2015) Cycadophila<\/em>, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis<\/em> (Cycadaceae) in Asia. Zootaxa<\/em>, 3986 (3), 251\u2013278.<\/p>

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Yoshida, T. & Leschen, R.A.B. (2020) Larval descriptions and exuvial retention of Toramini (Coleoptera: Erotylidae: Cryptophilinae). The Coleopterists Bulletin<\/em>, 74 (1), 1\u201314.<\/p>

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<\/p>"}],"container-title":["Zootaxa"],"link":[{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4838.2.7\/41013","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4838.2.7\/41013","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2023,2,23]],"date-time":"2023-02-23T01:47:56Z","timestamp":1677116876000},"score":17.867163,"resource":{"primary":{"URL":"https:\/\/www.mapress.com\/zt\/article\/view\/zootaxa.4838.2.7"}},"issued":{"date-parts":[[2020,8,26]]},"references-count":1,"journal-issue":{"issue":"2","published-online":{"date-parts":[[2020,8,26]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.4838.2.7","ISSN":["1175-5334","1175-5326"],"issn-type":[{"type":"electronic","value":"1175-5334"},{"type":"print","value":"1175-5326"}],"published":{"date-parts":[[2020,8,26]]}},{"indexed":{"date-parts":[[2024,3,18]],"date-time":"2024-03-18T01:12:26Z","timestamp":1710724346693},"reference-count":0,"publisher":"Magnolia Press","issue":"3","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"Asian species assigned to Pharaxonotha Reitter formed a separate clade from New World species of the genus and are distinguished on the basis of morphology and analysis of 16S rRNA gene sequences. Cycadophila, new genus, is described based on a consideration of four Asian species in comparison with New World Pharaxonotha. This new genus includes two new species (Cycadophila debaonica new species and Cycadophila fupingensis new species) and two previously described species (Cycadophila nigra (Gorham) new combination and Cycadophila yunnanensis (Grouvelle) new combination). These latter two may represent species complexes. Lectotypes are designated for Thallis nigra Gorham and Pharaxonotha yunnanensis Grouvelle. Larvae of C. debaonica and C. fupingensis are described based on individuals collected on the cones of Cycas debaoensis Y. C. Zhong & C. J. Chen (Cycadaceae), a cycad native to Guangxi and Yunnan provinces, China.\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.3986.3.1","type":"journal-article","created":{"date-parts":[[2015,7,17]],"date-time":"2015-07-17T00:59:10Z","timestamp":1437094750000},"source":"Crossref","is-referenced-by-count":7,"title":["Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae)\u00a0inhabiting Cycas debaoensis (Cycadaceae) in Asia"],"prefix":"10.11646","volume":"3986","author":[{"given":"GUANG","family":"XU","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"WILLIAM","family":"TANG","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"PAUL","family":"SKELLEY","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"NIAN","family":"LIU","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"STEPHEN","family":"RICH","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2015,7,17]]},"container-title":["Zootaxa"],"link":[{"URL":"https:\/\/mapress.com\/zt\/article\/download\/zootaxa.3986.3.1\/35697","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/mapress.com\/zt\/article\/download\/zootaxa.3986.3.1\/35697","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,3,18]],"date-time":"2024-03-18T00:37:05Z","timestamp":1710722225000},"score":16.031078,"resource":{"primary":{"URL":"https:\/\/mapress.com\/zt\/article\/view\/zootaxa.3986.3.1"}},"issued":{"date-parts":[[2015,7,17]]},"references-count":0,"journal-issue":{"issue":"3","published-online":{"date-parts":[[2015,7,17]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.3986.3.1","ISSN":["1175-5334","1175-5326"],"issn-type":[{"value":"1175-5334","type":"electronic"},{"value":"1175-5326","type":"print"}],"published":{"date-parts":[[2015,7,17]]}},{"indexed":{"date-parts":[[2025,12,11]],"date-time":"2025-12-11T20:49:10Z","timestamp":1765486150828},"reference-count":1,"publisher":"Magnolia Press","issue":"3","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"A new species of Erotylidae, Cycadophila (s. str.) mumia sp. nov., the first fossil representative of the subfamily Pharaxo-nothinae, is described and illustrated from Eocene Baltic amber. A second closely related specimen is reported but not described as new due to poor preservation in amber. The occurrence of representatives of the Recent cycad-specialized genus in Baltic amber can indirectly indicate the presence of their host plants (Cycadales: Cycadophyta: Cycadopsida) in the Eocene amber forest.\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.4337.3.6","type":"journal-article","created":{"date-parts":[[2017,10,19]],"date-time":"2017-10-19T00:56:35Z","timestamp":1508374595000},"source":"Crossref","is-referenced-by-count":9,"title":["First fossil representatives of Pharaxonothinae Crowson (Coleoptera:\u00a0Erotylidae): indirect evidence for cycads existence in Baltic amber forest"],"prefix":"10.11646","volume":"4337","author":[{"given":"VITALII I.","family":"ALEKSEEV","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"ANDRIS","family":"BUKEJS","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2017,10,19]]},"reference":[{"key":"41416","unstructured":"

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Valencia-Montoya, W.A., Tuberquia, D., Guzm\u00e1n, P.A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita<\/em> A. Lindstr. & Id\u00e1rraga by Pharaxonotha<\/em> beetles in the Magdalena Medio Valley, Colombia: a mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interaction<\/em>, 1\u201313. [published online]
https:\/\/doi.org\/10.1007\/s11829-017-9511-y<\/p>

Vovides, A.P. (1991) Insect symbionts of some Mexican cycads in their natural habitat. Biotropica<\/em>, 23, 102\u2013104.
https:\/\/doi.org\/10.2307\/2388697<\/p>

W\u0119grzynowicz, P. (2002) Morphology, phylogeny and classification of the family Erotylidae based on adult characters (Coleoptera: Cucujoidea). Genus<\/em>, 13, 435\u2013504.<\/p>

Xu, G., Tang, W., Skelley, P., Liu, N. & Rich, S. (2015) Cycadophila<\/em>, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis<\/em> (Cycadaceae) in Asia. Zootaxa<\/em>, 3986 (3), 251\u2013278.
https:\/\/doi.org\/10.11646\/zootaxa.3986.3.1<\/p>

<\/p>

<\/p>

<\/p>"}],"container-title":["Zootaxa"],"link":[{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4337.3.6\/12661","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4337.3.6\/12661","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2023,7,3]],"date-time":"2023-07-03T00:51:16Z","timestamp":1688345476000},"score":15.596956,"resource":{"primary":{"URL":"https:\/\/www.mapress.com\/zt\/article\/view\/zootaxa.4337.3.6"}},"issued":{"date-parts":[[2017,10,19]]},"references-count":1,"journal-issue":{"issue":"3","published-online":{"date-parts":[[2017,10,19]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.4337.3.6","ISSN":["1175-5334","1175-5326"],"issn-type":[{"value":"1175-5334","type":"electronic"},{"value":"1175-5326","type":"print"}],"published":{"date-parts":[[2017,10,19]]}},{"indexed":{"date-parts":[[2025,11,4]],"date-time":"2025-11-04T23:28:16Z","timestamp":1762298896749},"reference-count":1,"publisher":"Magnolia Press","issue":"2","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"Ceratophila new genus (Erotylidae: Pharaxonothinae) inhabiting male cones of the Mesoamerican cycad genus Ceratozamia Brongniart (Zamiaceae) is described for seven new species and compared to other genera of the Pharaxonothinae. Ceratophila differs from other genera of the Pharaxonothinae in many characters, most notable of which are: lateral pronotal carina thicker anteriorly in lateral view; head with supraocular and transverse occipital lines; elytra lacking a basal bead; and stridulatory files at base of head narrowly separated. Ceratophila has the male genitalia laterally compressed with a short median lobe and flagellum, similar to other Erotylidae. In comparison, the most superficially similar pharaxonothine genus Pharaxonotha Reitter has: lateral pronotal carina thin along entire length; head lacking supraocular lines, but with transverse occipital lines; elytra with distinct basal bead; stridulatory files at base of head widely separated; and unique male genitalia that are dorsoventrally flattened with a long median lobe and flagellum. Ceratophila contains two subgenera, which differ most notably in external characters and sexual dimorphism. Members of Ceratophila (Ceratophila), the nominal subgenus, have: more cylindrical bodies, pronotum evenly convex dorsally; the pronotum lacking basal longitudinal grooves on disc; metatibiae that are distinctly triangularly dilated toward apex; and no apparent external sexual dimorphism. In comparison, members of Ceratophila (Vovidesa) new subgenus have: dorsoventrally flattened bodies; the pronotum explanate (concave) along lateral margins; the pronotum with basal longitudinal grooves on disc; metatibiae weakly or not dilated toward apex; and with all tibiae in males showing strong sexual dimorphism. Four new species are described within the subgenus Ceratophila: C. (C.) chemnicki, C. (C.) gregoryi, C. (C.) picipennis, C. (C.) sanchezae; and three new species are described within the subgenus Vovidesa: C. (V.) chiapensis, C. (V.) mixeorum, C. (V.) vazquezi. Keys to genera, subgenera and species of known adults based on morphology are provided.\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.4508.2.1","type":"journal-article","created":{"date-parts":[[2018,10,31]],"date-time":"2018-10-31T00:23:30Z","timestamp":1540945410000},"source":"Crossref","is-referenced-by-count":8,"title":["Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)"],"prefix":"10.11646","volume":"4508","author":[{"given":"WILLIAM","family":"TANG","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"PAUL","family":"SKELLEY","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"MIGUEL ANGEL","family":"P\u00c9REZ-FARRERA","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2018,10,31]]},"reference":[{"key":"43884","unstructured":"

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<\/p>

<\/p>"}],"container-title":["Zootaxa"],"link":[{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4508.2.1\/16423","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4508.2.1\/16423","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2023,5,11]],"date-time":"2023-05-11T02:47:49Z","timestamp":1683773269000},"score":15.382664,"resource":{"primary":{"URL":"https:\/\/www.mapress.com\/zt\/article\/view\/zootaxa.4508.2.1"}},"issued":{"date-parts":[[2018,10,31]]},"references-count":1,"journal-issue":{"issue":"2","published-online":{"date-parts":[[2018,10,31]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.4508.2.1","ISSN":["1175-5334","1175-5326"],"issn-type":[{"value":"1175-5334","type":"electronic"},{"value":"1175-5326","type":"print"}],"published":{"date-parts":[[2018,10,31]]}},{"indexed":{"date-parts":[[2025,5,14]],"date-time":"2025-05-14T03:44:38Z","timestamp":1747194278935,"version":"3.40.5"},"reference-count":2,"publisher":"Magnolia Press","issue":"3","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"Two beetles, Ceratophila (Vovidesa) chipjonesi, new species, and Pharaxonotha perezi, new species, (Coleoptera: Erotylidae: Pharaxonothinae) are described from the male cones of the New World cycad Ceratozamia santillanii P\u00e9rez-Farr. & Vovides (Cycadales: Zamiaceae). Morphological analysis and keys are provided for distinguishing these from related beetles inhabiting Ceratozamia in Mexico.\r\n\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.5150.3.7","type":"journal-article","created":{"date-parts":[[2022,6,7]],"date-time":"2022-06-07T23:35:13Z","timestamp":1654644913000},"page":"428-442","source":"Crossref","is-referenced-by-count":1,"title":["Two new species of Pharaxonothinae beetles (Coleoptera: Erotylidae) inhabiting cones of the cycad Ceratozamia santillanii P\u00e9rez-Farr. &amp; Vovides (Cycadales: Zamiaceae) in Mexico"],"prefix":"10.11646","volume":"5150","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-9009-3113","authenticated-orcid":false,"given":"WILLIAM","family":"TANG","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7125-4639","authenticated-orcid":false,"given":"HECTOR GOMEZ","family":"DOMINGUEZ","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2022,6,8]]},"reference":[{"key":"120423","unstructured":"

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